BRITISH JOURNAL OF ENTOMOLOGY AND NATURAL HISTORY 



Editor: 



Richard A. Jones, B.Sc, F.R.E.S., F.L.S. 

 13 Bellwood Road 



Nunhead 

 London SE15 3DE 

 (Tel: 071 732 2440) 

 (Fax: 071 277 8725) 



Editorial Committee: 



Rev. D. J. L. Agassiz, M.A. T. G. Howarth, B.E.M., F.R.E.S. 



R. D. G. Barrington, B.Sc. I. F. G. McLean, Ph.D., F.R.E.S. 



E. S. Bradford Mrs F. M. Murphy, B.Sc. 



P. J. Chandler, B.Sc, F.R.E.S. M. J. Simmons, M.Sc. 



B. Goater, B.Sc, M.I.Biol. T. R. E. Southwood, K.B., D.Sc, F.R.E.S. 



A. J. Halstead, M.Sc. R. W. J. Uffen, M.Sc, F.R.E.S. 



R. D. Hawkins B. K. West, B.Ed. 

 P. J. Hodae 



British Journal of Entomology and Natural History is published by the British 

 Entomological and Natural History Society, Dinton Pastures Country Park, Davis Street, 

 Hurst, Reading, Berkshire RG10 OTH, UK. Tel: 0734-321402. 

 The Journal is distributed free to BENHS members. 



©1994 British Entomological and Natural History Society. 



Typeset by Dobbie Typesetting Limited, Tavistock, Devon. 

 Printed in England by Henry Ling Ltd, Dorchester, Dorset. 



BRITISH ENTOMOLOGICAL AND NATURAL HISTORY SOCDZTY 



Registered charity number: 213149 



Meetings of the Society are held regularly in London, at the rooms of the Royal 

 Entomological Society, 41 Queen's Gate, London SW7 and the well-known ANNUAL 

 EXHIBITION is planned for 22 October 1994 at Imperial College, London SW7. Frequent 

 Field Meetings are held at weekends in the summer. Visitors are welcome at all meetings. 

 The current Programme Card can be had on application to the Secretary, R. F. McCormick, 

 at the address given below. 



The Society maintains a library, and collections at its headquarters in Dinton Pastures, 

 which are open to members on the second and fourth Sundays of each month, telephone 

 0734-321402 for the latest meeting news. 



Applications for membership to the Membership Secretary: A. Godfrey, 10 Moorlea Drive, 

 Baildon, Shipley, West Yorkshire BD17 6QL. 



Subscriptions and changes of address to the Assistant Treasurer: M. G. Telfer, 12 Jasmine 

 Close, Norwich, Norfolk NR4 7NE. 



Non-arrival of the Journal, faulty copies or other problems arising from distribution of 

 the Journal or notices to the Distribution Secretary: D. Young, 9 Marten Place, Tilehurst, 

 Reading, Berkshire RG3 6FB. 



Orders for books and back numbers of the Journal and Proceedings to the Sales Secretary: 

 R. D. Hawkins, 30d Meadowcroft Close, Horley, Surrey RH6 9EL. 



General Enquiries to the Secretary: R. F. McCormick, 36 Paradise Road, Teignmouth, 

 Devon TQ14 8NR. Tel: 0626-779"543. 



Cover illustration: Carabid beetle. Photo: R. Williams. 



NOTE: The Editor invites submission of photographs for black and white reproduction 

 on the front covers of the journal. The subject matter is open, with an emphasis on aesthetic 

 value rather than scientific novelty. Submissions can be in the form of colour or black 

 and white prints or colour transparencies. 



