210 BR. J. ENT. NAT. HIST., 13: 2001 



the region involved should be counted as one segment showing incipient subdivision 

 or as two segments partially fused. Mason (1954) accepted the second interpretation 

 and appended to the total count two (or more) consecutive numbers joined by a plus 

 sign to denote the partially fused segments; this system, with the fused segments 

 shown in brackets, has been adopted here. Counts, expressed in this way, of flagellar 

 segments for nymphs are shown in Table 1, and for adults in Table 2. 



The differing total counts for any one of these later stages can be adjusted by the 

 addition of the first bracketed figure. Thus flagellar counts for third instar females 

 that, depending upon interpretation, could be regarded as ranging between 15 and 19 

 are brought to a common value of 22, a sum that increases by two at each subsequent 

 moult. This regularity has been of value in deducing the numerical stage when 

 rearing details were incomplete. In the tables, the number of individuals having any 

 one count is indicated by two figures separated by a colon. The first of these is the 

 number for which stage and developmental type are known from rearing history; the 

 second is the number assigned to that stage through a shared flagellar segmentation. 

 For the latter, inclusion in a particular developmental category has been based on a 

 combination of numerical stage thus deduced with a rearing history that, although 

 incomplete, includes stages N-l, N or adult. 



Growth of the hind femora 



The factorial increase in length of the hind-femora at each moult was calculated 

 for the entire sequence recorded for every individual of known or deduced nymphal 

 history. The combined values for years and sites are summarised for males and both 

 female sequences as ranges and means + one standard deviation of the mean (1 sem) 

 in Table 3. The relatively steady value of the factor over the successive nymphal 



Table 1. Flagellar segmentation of nymphal instars. 



Five-stage ? 

 1969 1970 



1976 



Four-stage $ 

 1969 1970 



1976 



Four-stage S 

 1969 



3rd instar 



16(6 + 7) 

 17(5 + 6) 



0:1 

 1:5 



0:1 

 0:2 



2:0 













18 (4 + 5) 



10:5 



2:9 



3:0 



1:2 



0:1 



3:0 



1:1 





19(3 + 4) 









1:1 



0:3 





4:9 



4th instar 



19(5 + 6) 



0:1 



0:1 















20(4 + 5) 



8:5 



1:33 



5:0 





0:1 









21 (3 + 4) 





0:1 





0:3 



0:3 



3:0 



4:4 



5th instar 



21 (5 + 6) 

 22(4 + 5) 

 23 (3 + 4) 



5:2 

 1:0 



0:1 



0:36 



0:1 



5:0 











Table 2. Flagellar segmentation 



of adults. 









Five-stage $ 

 1970 1976 



Four-stage $ 

 1969 1970 



1973 

 1975 



1976 



Four-stage 4 

 1976 



23 (3 + 4) 



24(4+5) 0:15 5:0 



0:1 0:1 



0:2 



3:0 



5:0 



