BLAKE.— ENCELIA AND RELATED GENERA. 353 
villous except for a submarginal naked border, narrowly white-mar- 
gined, awnless or with two stout subulate teeth or slender upwardly 
pubescent awns $-3 their length, connected by a fimbriate crown of 
nearly fused squamellae. 
Encelia (Geraea) nudicaulis Gray, Proc. Am. Acad. viii. 656 (1873); 
Jones, Proce. Calif. Acad. ser. 2. v. 701 (1895). 
Helianthella nudicaulis Gray, Proc. Am. Acad. xix. 9 (1883). 
Enceliopsis nudicaulis A. Nels. 1. c. (1909). 
Enceliopsis tuta A. Nels. 1. ec. (1909). 
Specimens examined: IpAuo: rather rare, dry rocky bluffs, Salmon 
River near Bay Horse, 5 Aug. 1895, Henderson 3653 (N); dry sage 
brush hills, above Salmon River, 6 Aug. 1895, Henderson 3653 (N); 
Nevapa: Candelaria, Esmeralda Co., 1881, Shockley (G); Hawthorne, 
Lepantha Mine, alt. 1677 m., 25 May 1897, Jones (N); compact clay 
slopes, alt. 305 m., Las Vegas, 29 April 1905, Jones 11857 (hb. Jones) ; 
limestone clays, Las Vegas, 4 May 1905, Goodding 2271 (G, type 
collection of E. tuta); clay, Horse Spring, alt. 915 m., 17 April 1894, 
Jones 5069k (hb. Jones); Uran: St. Thomas or St. George, Capt. 
F. M. Bishop (HoLoTYPE in Gray Herb.); gravel at foot of precipitous 
slopes in very poor clay soil, Marysvale, alt. 1830 m., 4 June 1894, 
Jones 5376 (hb. Jones); Ferguson Spring, alt. 1920 m., 14 June 1900, 
Jones 6403 (hb. Jones); halfway station W. of Wa Wa, alt. 2135 m., 
15 May 1906, Jones 11856 (hb. Jones).— Inhabits rocky or hard clay 
knolls where the soil is very compact (Jones, in litt.). 
I am unable to separate satisfactorily E. tuta from the older E. 
nudicaulis. The type of the latter has medium-sized orbicular leaves, 
connected by the Shockley and Jones plants with the small subacute 
ones of E. tuta, while the Henderson plants, largest- and broadest- 
leaved of all, bear some smaller leaves identical in shape and tip with 
those of E. tuta, indicating that the latter represents only a starved 
phase of E. nudicaulis. The achenes appear to be rather variable in 
pubescence when young, and at maturity are strongly bidentate or 
with two awns of varying length, the longest that I have seen being 
about ¢ the length of the mature fruit, although when young they are 
often longer relative to the ovary. The squamellae, fairly distinct when 
young, become fused into a barely fimbriate crown on the ripe fruit. 
*— Heads larger, rays 2-4. Ps em. long; pubescence ee leaves 
ombic-ovate, acute. 
++ aeenpy puberulent or “ahasoaes rays 3.5-4.2 cm. long, 6-12 mm. . wide. 
E. GRANpIFLORA (Jones) A. Nels. “Stems very thick and 
a branched and very short, woody, densely covered with very 
