BY E. J. GODDARD 861 



from it, and displaced towards the ventral side by a greatl}' 

 distended blood-laden space 



(c) The ccelomic spaces have the same character as those of 

 Trocheta and other Herpobdellids. They are arranged in a 

 symmetrical fashion on each side, as mostly hollow sacs of an 

 elliptical oblong or circular shape, as seen in transverse sections. 

 They make their first appearance some little distance behind the 

 female genital aperture. Each sac is lined by an epithelium 

 consisting of cells highly granular in character, the layer having 

 a ragged and slightly disintegrated nature as regards its con- 

 stituent elements. The granular nature of the cytoplasm of 

 these cells is in all probability due to the presence of pigment 

 granules in each cell, the whole system of these spaces being 

 developed in connection with the botryoidal tissue as in Trocheta 

 and Nephelis. They extend throughout the postgenital portion 

 of the body as far back as the anal region, and reach their 

 maximum in the middle part of the animal. In the extreme 

 anterior region of their occurrence they appear as three hollow 

 sacs on either side, two immediately dorsal to the ovary and the 

 third directly ventral to that organ and on a horizontal level 

 with the base of the pharynx which, just in this region, passes 

 into the oesophagus. The cavities carry some metameric signifi- 

 cance, and communicate with the lateral bloodvessel in each 

 somite. In the first few somites in which they occur, they have 

 the same distribution as described above. Behind this they form 

 a linear series on each side parallel to the sides of the alimentary 

 canal, those of one side communicating with the other side by 

 continuations of the pouches dorsal to the alimentary canal, these 

 connections occurring in each somite. Eventually we find the 

 alimentary canal invested by a series of such spaces arranged so 

 as to give the impression of a complete circle of the same in the 

 region of their greatest development. Although these spaces 

 approach into close association with the ventral sinus, no direct 

 communications between these two forms of chambers were made. 



In the region of the intestine enormously distended sinuses are 

 found laden with blood, some reaching such a strong development 

 73 



