119 



The peculiar quadrangular anthers figured in that work seem somewhat 

 diagrammatic. 



fcecunda. — Anthers ovate, with parallel distinct cells (B.F1.) Anthers nearly ovate, opening by 

 parallel slits (Eucalyptographia). 



I cannot find the connective a double gland, as shown in that work. 



loxophleba (considered a separate species in B.FL). — Anthers small, with parallel distinct cells 

 (B.F1.). 



It is further stated that the filaments are usually dark-coloured in the dried 

 specimens. I find this in fcecunda also. 



If the above definitions be analysed it will be observed how little different 

 the anthers of the two species are. When one comes to what I usually call 

 "transit " forms, it becomes frequently perplexing to say to what species a certain 

 anther belongs. As a rule, the anthers of E. fcecunda are. smaller and paler than 

 those of E. incrassata, but this character has its exception. I have, therefore, 

 thought it best to submit a number of drawings of anthers of the two species, from 

 illustrative specimens, and I think the anthers assist in strengthening the evidence 

 afforded by other parts of the plants, which I advance to show that there is real 

 affinity between E. incrassata and E. fcecunda, and that indeed they run into each 

 other. 



2. E. odorata, Behr. 



E. fcecunda and E. odorata specifically resemble each other very closely, and 

 frequently the leaves and fruits are difficult to separate. At page 114 1 have 

 expressed the opinion that E. loxophleba, Benth., var. fruticosa, Benth., is really 

 E. odorata. 



As a very general rule Bentham's description of the anthers (" very small 

 with globular distinct cells ") holds good for E. odorata. At the same time I would 

 like to emphasise the point that anthers vary like every other organ in Eucalyptus, 

 that they vary in size and also in shape. I have never seen them quite of the shape 

 of those figured by Mueller in Eucalytographia,* but certainly the openings do some- 

 times tend to depart from the circular form (pores) and tend to parallel openings. 



The timbers of E. fcecunda and E. odorata resemble each other a good deal. 

 But the two species are sharply separated by their juvenile foliage, that of 

 E. fcecunda being broadish, as figured (Plate 24) that of E. odorata being narrow, 

 oblong, and often with a mucrone. 



8. E. fruticetorum, F.v.M., Fragm. ii, 57. 



This is (as regards the Western Australian specimens) identical with 

 E. loxophleba, Benth. (B.F1. iii, 252.) I have shown (Part III, p. 80 of this 

 work) that E. fruticetorum is a synonym of E. calyco'jona, Turcz. 



* As figured in Eucalyptographia they are nearly square in outline ; the openings, as shown, are certainly not pores 

 and not parallel cells, but intermediate between ParallelantheraB and Renantherae. 



