468 



(b) Calyx-tufce. 



Historical. 



Fleshiness and other characters. 



Colour. 



Shape. 



Sculpture — 



(a) Angles and ribs. 



(5) Ribbing confined to the calyx-tube. 



Calyx-tube. — Obconical, campanulate or oblong, adnate to the ovary at the base or rarely to the 

 top, truncate and entire after the falling off of the operculum or with four minute teeth. (B. Fl. iii, 185). 

 The form and dimensions of the calyx-tube (hypanthium of Schauer, cupula of De Candolle) are taken 

 when the stamens are expanded but still adhering ; after they fall, it often alters so much that it neither 

 indicates the form it had in flower nor yet that which it will assume in fruit, (ib. p. 187). 



(The reference to the " 4 minute teeth " more particularly applies to the 

 Eudesmiea?, but in E. tetraptera (and to a less extent E. Forrestiana), there some is 

 approach to a 4-toothed calyx, and therefore affinity to the Eudesmieae.) 



Calyx-tubes usually free." — The tendency of the alabastra (buds) to become concrete (united in 

 growth) is well shown in such species as E. ruclis and E. obliqua, in which species they often (?) form dense, 

 spherical masses, their contiguous surfaces being quite flattened, so that the lower portions of the mature 

 seed-vessels assume the form of an inverted hexagonal pyramid, but actual concretion of these organs 

 never occurs in either of these species. The extraordinary form E. (SympJiyomyrtus) Lehmanni, offers a 

 remarkable instance of the concretion of the alabastra (buds) into a hard, woody mass, though each separate 

 flower still retains its individuality, and from the consideration of this fact, we are enabled to assert its 

 immediate derivation from E. cornuta, by the simple concretion of the alabastra. In this genus, and 

 indeed throughout the vegetable series generally, as compared with the higher animal orders, there seem 

 to have been preserved a vast number of transition forms, a fact that may be regarded as indicative of 

 vicissitudes in the conditions under which those forms exist rather than of any excess of susceptibility for 

 variation inherent in the forms themselves." (MSS. of Augustus Oldfield, about 1864.) 



As to E. Eehmanni—see Part XXXV, pp. Ill, 112, with Plate 144, Bentham 

 (B. Fl. iii, 233) refers to the calyx-tubes being more or less immersed in the globose 

 mass of the receptacle. 



Lateral connation of fruits is rare. Mr. A. D. Hardy figures (fig. 3, b, c, 

 Plate 13), Proc. Roy. Soc. Vict., xxix (New Series), 171, two cases in E. cordata. I 

 have seen a similar fusion of calyx-tubes in E. similis from Emerald, Queensland, 

 collected by R. Simmonds. 



Fleshiness and Other Characters. 



See under E. terminalis ; walls may be very thick, which dry in folds when not 

 ripe. Fleshiness is also notable in E. macrocarpa, E. pyriformis, E. tetraptera, and some 

 other species. 



We have a glandular calyx-tube in E, gracilis. See fig. D, Plate 12, Part III. 



In E. setosa, the calyx-tube is often more or less covered with bristles. See 

 fig. 6b, Plate 157. Part XXXVIII. 



