473 



Sculpture. 



(Angles and ribs, — raised markings.) 



Augustus Oldfield, Mueller's collector of over seventy years ago in Western 

 Australia and elsewhere, referred to this, and I quote from his MSS. about 1864 : — 



Among vegetables we find no less striking evidences of the derivation oi one species from another, 

 by the blending of organs, the generally superior size, and diminished number of such organs — besides the 

 evidence offered by the presence of ridges, grooves, and minute striae marking the points of concretion of 

 the blended individuals — invariably betraying their true origin. The genus Eucalyptus affords a remarkable 

 instance of this derivation of species, for starting from E. microtheca with its flowers in panicles containing 

 numerous individuals which are minute and semi-herbaceous, we may pass on by E . floribunda and E . rudis, 

 to E . <jo»i phocephal a and E. hypoleuca, and thence to such noble (sic) species as E. calopyhylla, E. Preissii, 

 E. crythrocorys and E. erythrocalyx, finally reaching that remarkable form E. tetraptera, the last of this 

 series, throughout which there is a gradual diminution in the number of alabastra (buds) in each aggregation 

 of the same, attended by a consequent increase in the dimensions of these organs. As we descend in this 

 series, we generally find the complexity of structure of the alabastra indicated by raised ridges (4-16). as 

 in E. erythrocorys, E. Preissii, E. tetraptera, E. erythrocalyx, as well as in E. globulus and many other 

 species; by grooves (16-32), as in E. costata and E. pleurocarpa; or by an immense number of delicate 

 stria', as in E. mlophylla, E. rudis, &c, and the same thing is oftentimes manifest in the aggregation of 

 stamens into four bundles, as in E. crythrocorys, E. eudesmioides, &c, a condition carried a step further in 

 Eudesmia (a genus which ought certainly to be included in Eucalyptus) where the operculum (combined 

 petals and sepals) is quadripartite. 



The following are arranged according to the numbers of the angles, ridges, or 

 ribs. They vary — (a) in number, and in some species may be absent; and (b) in 

 their prominence. 



E. Mooreana. — With one or two angles. 



E. canaliculata. — One or two opposite sharp ridges. 



E. Bancrofti. — Usually -with two angles. 



E. Lane-Poolei. — Two slightly raised ribs. 



E. diptera. — With two wings. 



E. Griffithsii. — Two of the ribs broadened almost into wings. 



E. notabilis. — Two angles or ribs sometimes so prominent as to be winged. 



E. platypus. — Terete or with two to four more or less prominent ribs or angles. 



E. piriformis. — More or less prominent two-four ribbed or almost winged. 



E. goniocalyx. — Often two to four prominently angled. 



E. Shirleyi. — Three prominent ribs with two or three less prominent ones. 



E. Forrestiana, E. tetraptera. — Quadrangular. In the latter species the ribs, which 



are so wide as to be " wings," terminate in short teeth at the commissure. 

 E. Stowardi. — Five prominent ribs. 

 E. corrugata. — Six to eight very prominent ridges. 

 E. ptychocarpa. — About eight prominent ridges. 

 E. torquata. — Seven to ten ridges. 

 E. setosa. — Often slightly eight-ribbed. 

 E. miniata. — More or less prominently eight-angled. 



