600 



Numbers of the Valves. 



Bentham ignores them. Mueller gives numbers for 27 out of about 100 

 (" Eucalyptographia ") with, the following results (the number of species is given in 

 brackets) : — • 



3 valves (1); 3, rarely 2 or 4 (1); 3-4 (3); 3, rarely 4 (3) ; 3 or much oftener 4 

 (1); 3 or oftener 4, rarely 5 (1); 3 or 4, exceptionally 5 (2); 3, rarely 5 (1); 

 4 (2) ; 4 or sometimes 3 or 5 (1) ; 4, rarely 5 (2) ; 4, rarely 3 or 5 (2) ; 4 or 5, 

 rarely 3 (1); 4 or 5, rarely 6 (2); 4-6 (1); exceptionally 5 (1); 5 or some- 

 times 4 or 6 (1) ; 5-6 or rarely 4 (1). Could anything be more confusing ? 



I have often sat under a tree examining the fruiting branches I had broken off, 

 and found the variation in the number of valves such that I came to the conclusion 

 that they had no diagnostic value in many species. 



Anomalous Valves. 



E. megacarpa F.v.M. Figured at fig. 6c, Plate 78, Part XVIII. 



" . . . . the rim very convex and prominent, continuous with the thick, conical, obtuse, incurved 

 and prominent valves of the cajjsule." (B.F1. iii, 232.) 



"'. . . . with 5 or sometimes 4 or 6 thick, emersed, convergent valves. . . ." 

 (" Eucalyptographia.") 



The valves of this species are anomalous in being fleshy and incurved. The 

 valves of E. globulus are somewhat similar to E. megacarpa in that in both species the 

 accessory covering of the valves is simply an enlargement of the disc which expands 

 and becomes considerably thicker than the valves as the fruit or capsule develops. 

 See. Disc, Part LXI. 



E. Preissiana. Figured at fig. 3. Plate 78, Part XVIII. 



" Disc broad and concave, the ovary with as many protuberances in the centre as valves." 

 (B.FL iii, 233.) 



" Valves 5-6, rarely 4, short, deltoid, permanently connivent, not protruding, surrounded by as 

 many or twice as many depressed protuberances." ( :; Eucalyptographia.") 



This species is anomalous in that the fruits have spheroidal protuberances on 

 the depressed rim. They are figured as above, and, under Fruits, I will later submit 

 more detailed drawings. The protuberances are not always 12 in number; they may 

 be 10, and the alternate ones may be absorbed, leaving 6 or 5. Their morphological 

 significance will be explained in Part LXI. 



RIM. 



■ * 



No one appears to have defined the " rim," and indeed it is somewhat vague. 

 It is usually taken to include everything from the commisural line or where the calyx- 

 tube meets the operculum, to the base of the valves of the capsule where the capsule is 

 not much sunk. 



