1917] Butters,— Studies in Ferns — Athyrium 199 
variety sometimes 2 m. tall, pinnae 1-2 dm. long, remote, the lower 
ones with 5-10 cm. intervals; pinnules triangular-lanceolate, 1.5-3 cm. 
long, 0.5-1.5 em. wide, remote, separated by about their own width, 
incised with numerous low, broad and blunt teeth, or in the largest 
fronds, pinnatifid, the segments of the third order oblong, obtuse, 
slightly 5-8 toothed, the largest 6 X 2.5 mm. in size. 
This variety differs from A. Filiz-femina var. multidentatum (Déll) 
Milde of Europe, chiefly in its relatively broad and short ultimate 
segments with almost suppressed marginal teeth, which are crowded 
towards the very obtuse apex of the segment. The immature form of 
this variety is the forma Hillii.2, The pinnules at this stage are large 
(4-5 X 7-10 mm.), very blunt and rounded, crowded, and not at all 
pinnatifid, but with a somewhat crenate outline and numerous very 
small teeth borne chiefly at the summit of the pinnule. This type of 
foliage is retained until after fruiting commences, and until the fronds 
are 5-6 dm. tall. The transition to the mature form may sometimes 
has already been pointed out, there is nothing about the sori of the American ferns of this 
species to distinguish them ‘ap aehess: ra a ian, German, and British specimens. Ac- 
cordingly, Ruprecht’s definitio . evel m becomes nearly meaningless, and his refer- 
ence to Lapland in his Gates nas et anal that he was including in his variety all 
ol 
ceetalicn ; am following Milde, who says (Fil. Eu. et At. p. 50, 1867.) “var. cyc! 
Whewtohe (List of N. A. hepa 31) says, ‘“‘The only [specimens] in the U. S. which can 
msidered as something like type specimens are two in the Gray herbarium which were re- 
ceived _ St. Peer ga of theme came from the Amur region, the other from Unalaska 
he type loca loca ™T, 1 t be identified, as there are 
in the is | i f the Amur region, all received by way 
of Poleverad: and all labelled Aapidian | Filiz spiasbe None of these shows any especial 
resemblance tot the west American orms of this species. The Unalaska ator is labelled, 


While this is of etd some te: the original Apne pone eagle Ruprecht published ae ¥ 
name, nor any recogni: ample 
sitchense as here defined. A specimen labelled in the same hand-writin o, * Athyn: rium Filix 
foemina (L.) Roth. var.— Kamtschatka”’ is sag A. Filiz-femina. 
The synonymy of the var. sitchense is, therefore 
ArayriuM Fiirx-remina (L.) Roth. var. sItcHENSE Ruprecht ex Moore, Index Fil. 
183 (1860). 
5 Athyrium sttchense Rupr. Dist. Crypt. Vas. Ross., 41 (1845). 
part. 
2 ATHYRIUM Finix-remina (L.) Roth. var. acnccesian Ruprecht ex Moore forma 
(Gilbert) eee nov., Athyrium cyclosorum f. Hillii Gilbert, List of N. A. Pterid, 32 (1901). 
