268 ° ROBINSON. 
included even those of wide distribution and in some cases of weedy 
character) somewhat exceeds the element common to Ecuador (19 
species). On-the other hand the actual affinities of the Colombian 
Eupatoriums, when determined by the more characteristic and more 
certainly native species of limited range, are more strongly Ecuadorian 
than Venezuelan. 
There seems no reason to suppose that the statistical relations here 
presented on the basis of this large and diversified genus will not be 
found to hold their approximate ratio in many other groups. Un- 
fortunately the present state of knowledge regarding the details of 


plant-distribution in Colombia, as well as of the geological, meteoro- _ 
logical, and physiographic conditions, is still much too slight to permit 
generalization regarding ecological relations or soil-influence in de- 
termining the present flora. 
EUPATORIUM [Tourn.] L. Heads homogamous, (1-4)5-«-flow- 
ered. Involucre cylindrical to campanulate; the scales of indefinite 
number, usually numerous, commonly graduated and appearing to be 
arranged in 2-8 series, the inner progressively longer, more rarely almost 
all subequal and with only a few (1-3) of the outermost considerably 
shorter. Receptacle flat or in varying degree convex to conical, 
glabrous or in one section pubescent, calvous. Corolla tubular, with 
or without a perceptibly enlarged campanulate, turbinate, or cylindn- 
cal throat; the limb (4—)5-toothed, regular. Anthers mostly connate 
but sometimes nearly or quite free, rounded (to rarely and obscurely 
cordate or subsagittate) at the base, provided at the summit with an 
ovate or oblong mostly obtuse rarely retuse membranaceous append- 
age. Style-branches much exserted, the appendage elongate, filiform 
or more commonly somewhat clavate, smooth or microscopically 
papillose. Achenes 5-angled, the angles usually rib-thickened and of 
modified texture, color, or pubescence, the intervals flat or often 
scope) hispidulous, never strongly plumose.— Sp. Pl. ii. 836 
L. f. Suppl. 354 (1781); HBK. Nov. Gen. et Spec. iv. 105-134 (1820), 
excl. certain species; DC. Prod. v. 141-186 (1836); Benth. & Hook. 
f. Gen. ii. 245 (1873); Bak. in Mart. FI. Bras. vi. pt. 2, 274-365, t. 
76-96 (1876); Klatt in Engl. Bot. Jahrb. viii. 33-36 (1887); Hoffm. 
’ in Engl. & Prantl, Nat. Pflanzenf. iv, Abt. 5, 138 (1890); Hieron. 
(1753); 
