42 Contributions from the Gray Herbarium 
involucre and should be kept distinct. The above interpretation 
_ of Actinea necessitates the following new combinations. 
omb. 
nov. Actinella depressa T. & G., var. pygmaea Gray, Mem. Am. 
Acad. ser. ii. iv. 100 (1849). Tetraneuris depressa (Gray) Woot. 
& Standl. Contrib. U. S. Nat. Herb. xvi. 193 (1913). 
This plant is confined to New Mexico and its leaves are glabrate 
or more or less silvery with an appressed silky pubescence. The 
leaves of typical A. depressa (T. & G.) Ktze., so far known only | 
from Utah, are green and glabrous or sparsely long-villous. 
“ ACTINEA ACAULIS (Pursh) Spreng., var. lanata (Nutt.), comb. 
nov. Actinella lanata Nutt. Trans. Am. Phil. Soc. ser. ii. vii. 379 
(1841). Tetraneuris lanigera Daniels, Univ. Mo. Stud. Sci. ii. 393 
(1911). A. lanigera Daniels, 1. ¢. 
The variety differs only in the looser pubescence and a some- 
what different habit. Dr. Gray included this plant in A. acaulis 
(Pursh) Spreng. but it may be recognized as a geographical variety. 
A. acaulis in typical form is essentially a species of the Great 
Plains and is characterized by a closely appressed silvery pubes- 
‘cence. Its scapes are usually 1-2 dm. high. The var. lanaia on 
the other hand is typically of the high mountains, is loosely vil- 
lous and infrequently raises its scapes to a height of 1 dm. But 
plants intermediate in character between the typical form and the 
variety are not infrequent, as, for instance, Murdoch’s no. 4042 
from the Black Hills of South Dakota and Sturgis’s material from 
Colorado Springs. The var. lanata occurs in two forms represent- 
ing extreme development in opposite directions. One form grows 
on the high plains in rocks or on dry gravel bars and has reduced 
leaves and heads and a densely caespitose habit; the other form 
is a tall state with elongate scapes and except for the lanate pube- 
scence closely approaches the glabrate variety of A. acaulis. These 
forms of the variety lanata may be known as 
~ ACTINEA ACAULIS (Pursh) Spreng., var. LANATA (Nutt.) Macbr., 
forma caespitosa (A. Nels.), comb. nov. Tetraneuris acaults 
(Pursh) Greene, var. caespitosa A. Nels. Bot. Gaz. xxviii. 127 (1899). 
T. brevifolia Greene, Pitt. iii. 266 (1898). 
“ACTINEA ACAULIS (Pursh) Spreng., var. LANATA (Nutt.) Macbr., 
forma arizonica (Greene), comb. nov. Tetraneuris coteage 
oy Pitt. iii. 266 (1898), not Rydberg, N. A. Fl. xxxiv- 
