TROPICAL AMERICAN COMPOSITAE, _ 35 
Santander, Colombia, 16 Jan. 1918. These correspond closely in all 
essential and most minor features with the original material of E. 
Squiresii from the delta of the Orinoco and certainly appear to be 
conspecific with it. The only differences found during a rather 
detailed examination were that the leaves of the Colombian plant 
were slightly firmer in texture and even more shortly petioled or 
subsessile. These differences are precisely of the kind and degree 
that occur very frequently between exposed and shade forms of 
the same plant. The range of the species is thus extended some 
1500 km. and over the watershed from the Orinoco Valley into 
that of the Magdalena. However, there is little difference in the 
latitude, and the habitat, low alluvial thickets, is similar. Renewed 
examination of the plants of this group brings out what had not 
been previously noticed, namely, the affinity between this species 
and E. turbacense Hieron. Of the latter species the writer has not 
seen the type, Stiibel’s no. 51, collected at Turbaco, Dept. Bolivar, 
in the delta region of the Magdalena. However, Hieronymus when 
publishing upon the plants of Lehmann (Engl. Bot. Jahrb. xxviii. 573) 
identifies with his E. turbacense Lehmann’s no. 5971, and of this a 
leaf and a bit of the inflorescence were received at the Gray Herbarium 
Some years ago in an exchange from the Royal Botanical Garden at 
Berlin. This plant of Lehmann’s was collected on the Rio Ortega 
in the Dept. Cauca, that is to say, some 900 km. to the south of the 
original station. If Hieronymus has been right in referring it to his 
E. turbacense, the following differences may be pointed out between 
that species and the later E. Squiresit. In E. turbacense the leaves 
are entirely glabrous above, while in E. Squiresti they are puberulent 
at least on the midnerve and sometimes perceptibly so on the surface 
as well; in E. turbacense the lowest two pairs of lateral veins leave the 
midnerve at an angle of about 40° and in length considerably exceed 
those arising at a greater distance from the base, while in EF. Squiresti 
the lower pairs of veins are no longer, indeed are usually shorter than 
Some of the others, and all of them leave the midnerve at a considerable 
angle, usually at about 70°. In E. turbacense the involucre is cam- 
Panulate, in E. Squiresii it is somewhat longer and campanulate- 
subeylindric (in the fresh state) or campanulate-subturbinate (in ~ 
the dried state). In E. turbacense the pubescence of the pedicels is 
distinctly longer and more sordid-tawny than in E. Squiresii. The 
© Species are certainly very close. In both the lower leaves have a 
Peculiar form, the lance-oblong blade being narrowed at the base into 
@ more or less elongated portion like a broadly winged petiole. It 
