Marsh Collection, Peabody Museum. 429 



down to its attachment upon the base of the second meta- 

 tarsal, and the groove is relatively large ; whereas in Dromo- 

 cyon it is small, and the tendon passed down to the outer 

 edge of the tubercle, thence turned abruptly inwards to be 

 inserted in the same place. The relatively large size of the 

 peroneal tubercle, which serves principally for the attachment 

 of the calcaneo-cuboid ligament, may be taken to indicate 

 unusual firmness of the tarsal joint. 



The navicular has much the same general shape as in / the 

 dog, and differs from it in that it is narrower, and its proximal 

 surface more deeply saddle-shaped ; it articulates with the 

 cuboid in quite the same way, and distally exhibits the three 

 usual facets for the cuneiformia. Posteriorly it presents 

 some important differences from that of the dog. In the 

 case of the latter, there are two more or less distinct navicu- 

 lar tubercles, separated by a wide sulcus. In Bromocyon 

 there is a single tubercle, which is produced into a long 

 pointed process. The three cuneiformia differ from those of 

 the dog, in that they are decidedly higher and narrower from 

 side to side ; their manner of articulation is, however, much 

 the same. The internal cuneiform is large and elongate, 

 bearing upon its distal extremity an articular facet for a rudi- 

 ment of the first metatarsal. 



The metatarsals closely resemble those of the dog, not only 

 in their elongate, compressed, and highly interlocking charac- 

 ter, but they show a decided tendency towards lateral flatten- 

 ing of their contiguous surfaces. Their distal ends, moreover, 

 exhibit that characteristic " square-cut " appearance so com- 

 mon to the modern Canidse. The foot is of the strictly 

 paraxonic type, the third and fourth metapodials being of 

 equal length. While the second and fifth are also of equal 

 length, the second is much the larger and stouter bone, the 

 disparity in size being considerably greater than that seen in 

 the dog, and about equal to that of the fore foot. The distal 

 metapodial keels are present and well developed ; they are, 

 however, confined to the plantar surface and do not extend 

 more than half-way around upon the dorsal surface. 



The proximal phalanges are proportionately shorter, 

 stouter, and more robust than in the dog, but the articular 

 faces have very much the same disposition and extent. 

 Proximally the plantar notch is not so deeply incised as in 

 the phalanges of the dog, but their dorso-plantar curvature 

 appears to be somewhat greater. The median phalanges, or 

 those of the middle row, are proportionately still shorter than 

 those of the first, as compared with the dog. They exhibit a 

 slight degree of distal asymmetry. The unguals are short, 

 rather broad, and deeply fissured. 



