80 T. Holm — Anemiopsis Calif ornica. 



from the ventral face and but slightly differentiated on that of 

 the dorsal. The homogeneous, somewhat open mesophyll con- 

 tains a little chlorophyll, and about seven very small mestome- 

 bundles are located in the middle of this tissue. 



The stem. 



The long stolons above ground and the flower-bearing stems 

 show the same structure. They are nearly cylindric, slightly 

 hairy, but perfectly smooth. We notice, also, here a wrinkled 

 cuticle, covering an epidermis of relatively small cells of which 

 the outer wall is distinctly thickened ; none of the epidermal 

 cells were transformed into secreting cells. Underneath the 

 epidermis is a hypoderm of only one layer of very large, 

 roundish cells, much larger than those of the adjoining cortex. 

 This tissue, the cortex, consists of about fifteen strata, of 

 which only the peripheral contain chlorophyll ; the innermost 

 layer is differentiated as a thin-walled endodermis, surrounding 

 a continuous ring of about fifteen layers of very thick-walled 

 stereome. Directly bordering on the stereome is a circle of 

 twenty-four collateral mestome-bundles separated from each 

 other by broad medullary rays ; they are oval in cross-sections 

 and contain both leptome and hadrome, the latter consisting of 

 a few, but wide vessels. But there is no parenchyma-sheath 

 and no mechanical support on the sides of 'these mestome- 

 bundles or around the hadrome. The central pith is very thin- 

 walled and open. Secreting cells abound in the cortex and 

 in the pith. 



The rhizome. 



The horizontal rhizome is cylindric, glabrous and smooth ; 

 towards the apex it is densely covered with sheathing, green 

 leaves, and is not exposed to the light. Its free portion, 

 behind the rosette of leaves, becomes soon deprived of both 

 the epidermis and hypoderm, but protected by many layers of 

 cork. The cortical parenchyma consists of about fifteen layers 

 with wide intercellular spaces ; the cells are filled with starch 

 or contain a secretion of a reddish brown color. The very 

 numerous mestome-bundles are narrow oblong in transverse 

 sections, and are nearly all arranged in a circle separated from 

 each other by rays of the very broad, central pith, which con- 

 tains starch. The leptome occupies only a small portion of the 

 mestome-bundles, and between this and the very prominently 

 developed hadrome are several strata of cambium. 



The root. 



All the roots were so far advanced that they showed only a 

 secondary stage of growth. By the increase in thickness the 



