Marsh Collection, Peabocly Museum. 149 



common in the Bridger species. I give in this connection (Plate 

 VI) a drawing of a well-preserved skull of this species in the 

 Marsh Collection, from the White Kiver Bad Lands of Nebraska, 

 on account of the apparently important position which it holds 

 with reference to the origin of certain living species. It has 

 been suggested by Matthew and myself,* that Daphcenus 

 vetus was the direct forerunner of Temnocyon of the John 

 Day Miocene, which in turn passed by gradual transition into 

 the living Dohles, or Ked Dogs of India. If this chain of 

 ancestry is correct, the evidence in favor of which seems to 

 be very direct and positive, it establishes a long and eventful 

 history for this phylum. 



Daphcenus Dodgei Scott, on the other hand, may be 

 regarded as the successor of the short-jawed species of Uin- 

 tacyon, of which TJ. vorax is a good type, and here again the 

 characters, as far as we know them, fit with apparent accuracy. 

 Daphcenus Dodgei is known from a lower jaw only, and may, 

 as suggested by Scott,* prove to be a distinct genus. Further- 

 more, it will not be surprising to find that it had already 

 developed a reduced dentition, and was leading into the short- 

 jawed dogs of the John Day, such as Oligobunus, Enhydix)- 

 cyon, Hycenoeyon, and thence through an unknown type, 

 into the living Bush Dogs, Icticyon, of the South American 

 tropics. 



Prodaphcenus Wortman and Matthew. 



Prodaphcenus Wortman and Matthew. Bull. Amer. Mus. Nat. Hist., 1899, 

 p. 114. 



In this group I arrange all those short-jawed dogs of the 

 Eocene in which the size of the premolars, with the exception 

 of the fourth superior, is much reduced. The dental formula 

 is not fully known, but is very probably I. f , C. |- Pm. |-, M. f . 

 The canines are large and more or less laterally flattened, the 

 heel of the inferior sectorial is basin-shaped, the anterior 

 border of the superior molars extended transversely, the two 

 external cusps are unequal in size and the postero-internal 

 cingular cusp is small or absent. 



The genus was established upon an incomplete superior 

 dental series from the Uinta deposits and was unfortunately 

 named. At the time, it was thought to be related to Daphcenus, 

 but it is now perfectly evident that it belongs to another line. 

 The only other group of Canidae thus far known in which this 

 combination of characters occurs is the Amphicyon series, and 

 as this begins abruptly in Europe in beds corresponding 



* Loc. cit , p. 115. 



f Notes on the Canidae of the White Kiver Oligocene. Trans. Amer. Philos. 

 Soc., 1898. 



