Chap. XI. 



OP REPRODUCTION AND VARIATION. 



393 



footstalk, is green, the other half being brown and of the nature of the 

 pomrne gris apple, with the line of separation between the two halves 

 exactly denned. The tree was a grafted one, and Mr. La Tonche thinks 

 that the branch which bore this curious apple sprung from the point 

 of junction of the graft and stock : had this fact been ascertained, the 

 case would probably haye come into the small class of graft-hybrids 

 presently to be given. But the branch may have sprung from the stock, 

 which no doubt was a seedling. 



Prof. H. Lecoq, who has made a great number of crosses between the 

 differently coloured varieties of Mirabilis jalapa™ finds that in the 

 seedlings the colours rarely combine, but form distinct stripes ; or half the 

 flower is of one colour and half of a different colour. Some varieties 

 regularly bear flowers striped with yellow, white, and red ; but plants of 

 such varieties occasionally produce on the same root branches with uni- 

 formly coloured flowers of all three tints, and other branches with half-and 

 half coloured flowers and others with marbled flowers. Gallesio 108 crossed 

 reciprocally white and red carnations, and the seedlings were striped; 

 but some of the striped plants also bore entirely white and entirely red 

 flowers. Some of these plants produced one year red flowers alone, and in 

 the following year striped flowers ; or conversely, some plants, after having 

 borne for two or three years striped flowers, would revert and bear exclu- 

 sively red flowers. It may be worth mentioning that I fertilised the Purple 

 Sweet-pea (Lathyrus odoratus) with pollen from the light-coloured Painted 

 Lady: seedlings raised from one and the same pod were not inter- 

 mediate in character, but perfectly resembled both parents. Later in 

 the summer, the plants which had at first borne flowers identical with 

 those of the Painted Lady, produced flowers streaked and blotched 

 with purple ; showing in these darker marks a tendency to reversion to 

 the mother- variety. Andrew Knight i09 fertilised two white grapes with 

 pollen of the Aleppo grape, which is darkly variegated both in its leaves 

 and fruit. The result was that the young seedlings were not at first 

 variegated, but all became variegated during the succeeding summer; 

 besides this, many produced on the same plant bunches of grapes which 

 were all black, or all white, or lead-coloured striped with white, or white 

 dotted with minute black stripes ; and grapes of all these shades could 

 frequently be found on the same footstalk. 



In most of these cases of crossed varieties, and in some of 

 the cases of crossed species, the colours proper to both parents 

 appeared in the seedlings, as soon as they first flowered, in the 

 form of stripes or larger segments, or as whole flowers or fruit 

 of two kinds borne on the same plant ; and in this case the 

 appearance of the two colours cannot strictly be said to be due 

 to reversion, but to some incapacity of fusion, leading to their 



107 • Geograph. Bot. de l'Europe/ "» 'Trait* du Citrus,' 1811, p. 48. 



torn, iii., 1854, p. 405 ; and ' De la 109 ' Transact. linn. Soc.,' vol. ix. p. 



Fecundation,' 1862, p. 302. 268. 



