f 



182 



HYBRIDISM. 



Chap. XIX. 



considerable interval of time, placed on the 



gly prepotent that it generally annihilates 



the effect of the foreign pollen 



with the pollen of the 



several forms of the same species, for legitimate pollen is strono-ly 

 prepotent over illegitimate pollen, when both are placed on the 



same stigma 



I ascertained 



by fertilisin 



al flow 



first illegitimately, and twenty-four hours afterwards legitimately 



with pollen taken fi 



the 



seedling 



peculiarly coloured variety, and 



i were similarly coloured; this shows that the 

 legitimate pollen, though applied twenty-four hours subsequently, 

 had wholly destroyed or prevented the action of the previously 



pplied illegitim 



pollen 



A 



6 



as, in making recipr 



crosses between the same two species, there is occasionally a 



result, so the same thing occurs with 



great 



differ 



the 



trimorphic plants ; for instance, the mid-styled form of Lytlirum 

 salicaria could be illegitimately fertilised with the greatest ease 

 by pollen from the longer stamens of the short-styled form, and 

 yielded many seeds ; but the latter form did not yield a single 

 seed when fertilised by the longer stamens of the mid-styled 

 form. 



In all these respects the forms of the same undoubted species, 

 when illegitimately united, behave in exactly the same manner 

 as do two distinct species when crossed. This led me carefully 

 to observe during four years many seedlings, raised from several 



illegitimate 



The chief result is that these illeg 



plants, as they may be called, are not fully fertile. It is pos- 

 sible to raise from dimorphic species, both long-styled and short 



tyled illegitimate plants, and from trimorphic plants all thi 



illegitimate forms. These can then be properly united in a 

 timate manner. When this is done, there is no apparent rej 

 why they should not yield as many seeds as did their par 



when legitimately fertilised. But such is not the 



they 



all infertile, but in various degrees ; some being so utterly 



incurably sterile that they did 



single 

 which 



yield during four 



seed or even seed-capsule. These illegitimate plants 

 ire so sterile, although united with each other in a legi- 



timate manner, may be strictly compared with hybrids when 



crossed inter 

 generally art 



and it is well known how 

 When, on the 



these latter 



other hand, a hybrid is crossed 



( 



] 



