May, Igo09.] THE ORCHID REVIEW. 131 
Neuwiedia looks more like some Amaryllidaceous plant than an Orchid, 
owing to its free stamens, elongated versatile anthers, and slender straight 
style. The perianth is nearly regular, and connivent into a narrow tube, 
which is inserted obliquely on the pedicel, while the lip is petal-like. Thus 
the flower owes its irregularity chiefly to the fact that three of the six 
stamens are absent, and these all on one side of the flower. The stamens 
present are the median one of the outer staminal whorl, and the lateral 
pair of the inner whorl. The flower is evidently fertilised by insects, 
though no details are known. In Apostasia, the next genus, there are only 
two stamens, the median one being reduced to a filament-like staminode, 
which is partly united to the back of the style, while the anthers are 
obliquely attached to a short filament and unequal at the base. The 
perianth is nearly regular, and the segments rather more spreading than in 
Neuwiedia. In Adactylus the perianth agrees with Apostasia, but the 
median stamen is entirely suppressed, and the anthers are cordate, and 
equal at the base. 
In Cypripediez a higher degree of specialisation is reached, but the plan 
of the flower is identical with that of Apostasiez. The lip has become 
transformed into a pouch-like organ, formed by the union of the median 
petal with a pair of petaloid staminodes, representing the two lateral 
stamens of the oufer staminal whorl (which are suppressed in Apostasiez) 
The lateral pair of the inner whorl are fertile, as in the preceding tribe, but 
have become globose in shape, and are borne upon a well-developed 
column, while the pollen grains cohere together in a sticky mass. The 
median stamen of the outer whorl is modified into a shield-shaped 
staminode, which is placed just behind the opening of the pouch, and the 
stigma is also shield-shaped, and situated on the underside of the column. 
The flowers, so far as known, are fertilised by bees, which enter the 
mouth of the pouch, crawl up the tube past the stigma, and make their exit 
at the lateral openings situated at either side of the base, opposite the 
anthers. In crawling out their backs become smeared with the sticky 
pollen, some of which is left on the stigma of the next flower visited. The 
whole flower is beautifully adapted to secure cross-fertilisation by suitable 
insects, and to exclude others whose visits would not secure this end. The 
insects are only known in a few cases, but without them seeds are not 
produced, and the inference is obvious. 
The tribe contains four genera, Selenipedilum, containing three tall 
reed-like plants, with small flowers, a three-celled ovary, a persistent 
perianth, and crustaceous seeds, as in Vanilla; Phragmopedilum, differing 
in its stemless habit, conduplicate leaves, larger flowers, deciduous 
perianth, and fusiform, reticulated seeds; Cypripedium, with one-celled ovary, 
deciduous habit, plicate leaves, and persistent perianth ; and Paphiopedilum, 
