WILLIAMS: THE GENUS DESMATODON 211 
Type LOcALITY: Rocky Mountains of British America. 
DisTRIBUTION: Greenland; Beechey Island, Arctic America; the 
Canadian Rockies; also in Europe. 
ExsIccATAE: Drummond, Musci Am. 145. 
ILLUSTRATION: B.S.G. Bryol. Eur. pl. 136. 
3. DESMATODON GUEPINI B.S.G. Bryol. Eur. (18-20): Desma- 
todon 8. 1843 
Trichostomum Guepini C. Miill. Syn. 1: 590. 1849. 
Barbula Guepini Schimp. Syn. ed. 2,197. 1876. 
Tortula Guepint Broth. in E. & P. Nat. Pfl. 13: 430. I902. 
Autoicous, the two or three very small male flowers scattered 
along the stem and composed of four or five pale, ecostate, ovate, 
- acute leaves, smooth or nearly so, the outer longer ones about 0.5 
mm. long, enclosing three or four antheridia about 0.25 mm. long, 
with few or no paraphyses: fertile plants rather loosely cespitose, 
bud-like, 1-3 mm. high; the larger upper leaves ovate to somewhat 
spatulate, with blade 1-1.5 mm. long, the apex somewhat rounded 
or acute, the margins entire and revolute from near the apex al- 
most to base and the costa mostly smooth on the back, excurrent 
into a nearly smooth point one fifth to one half the length of the 
blade; costa in cross-section showing mostly two guide-cells, four 
or five cells of about the same size in one row on the ventral side 
and on the dorsal side a thick stereid band with outer cells differ- 
entiated; leaf-cells rather obscure and densely papillose in upper 
part of leaf, more or less four to six sided, not or scarcely elongate, 
14-16 » wide, those of basal part smooth, pale, larger, more or less 
rectangular: outer perichaetial leaves not differentiated, the inner 
small, acutely pointed, with flat margins; seta erect, about 8 mm. 
ong; capsule erect, oblong-cylindric, 1-1.5 mm. long without 
lid, the stomata in one row near the base; annulus narrow, per- 
sistent, of one or two rows of cells; peristome pale, densely papil- 
lose, of sixteen slightly oblique, narrow teeth mostly divided nearly 
to the base into two filiform forks from a basilar membrane ex- 
tending well above the annulus; lid high-conic, its height about 
twice the basal diameter, the cells a little above the base elongate 
in nearly erect rows; spores nearly smooth, the larger 16 in diam- 
eter; calyptra cucullate, descending about half way down the cap- 
sule. [Fic. 3.] 
TYPE LOCALITY: France. 
Distrisution: California and France. 
ILLustRATION: B.S.G. Bryol. Eur. pl. 133. 
