A. B. Srour 101 
formed closely to the original description (Hemsley, 1905) of N. Forgetiana, 
but these were also plants whose flowers were of lighter and of darker 
shades of red, and there were green-flowered and white-flowered plants. 
It may be noted here that no pure stock of this species was ever 
kept in culture from the original and only importation of seed obtained 
from a wild source. In reply to inquiry, the author learns the following 
by letter from the firm of Sanders of St Albans, England, who made 
the importation : “We never sold any of the original stock. We simply 
kept them for hybridizing purposes, afterwards selling the entire lot 
together with the seed. We had previously secured a batch of really 
fine new crosses with NV. affinis, ete. resulting in a lovely set. . When 
these plants were in full bloom in all the shades of rose, purple, white, 
pink and crimson the effect was magnificent. We did not continue 
with the growing of the Nicotianas but sor our stock, We regret very 
much having lost this remarkable species.” 
Series 2. A self-fertile plant of the 1917 crop which appeared to be 
identical with the original description of V. Forgetiana was the parent of 
24 plants grown under field conditions. Guarded self-pollinations were 
made by hand at frequent dates from the first to the very last day of 
bloom. 
Eleven plants were entirely self-incompatible. They bloomed for 
periods of from 22 to 65 days, and the total of all flowers selfed was 377. 
One plant was judged as fully self-compatible. It produced fine 
pods in all flowers selfed on the first, second and fifth days of bloom. 
No controlled selfings were made thereafter. 
The other 12 plants were partially or feebly self-compatible. The 
complete data for these would occupy much space, but as the results 
have a very special bearing on the variations in self-fertility they can 
be summarized, at least for several plants which are typical for the 
results.’ 
One plant began blooming on July 27 and was still bane on 
Noy. 6 when there was a heavy frost. A total of 80 flowers was selfed 
on twenty different dates. On the third day of bloom the fourth flower. 
to open produced a pod with seeds ; 2 pods developed from the 2 flowers 
- that opened on the fourth day; one of 2 flowers selfed on the twelfth 
day developed a a pod; 3’selfed on the seventeenth day gave pods; on 
Sept. 4th, 6 out of 10 flowers selfed gave pods, but on ten dates im- 
mediately preceding, self-pollinations of 27 flowers all failed. From 
Sept. 4 until the end of the season about half of the flowers selfed 
produced pods and seeds. In respect to the proportion of flowers which 
Journ. of Gen. 1x 7 
