24 



PENNSYLFANIAN SPORES OF ILLINOIS 



radii are slightly concave and the corners 

 are slightly rounded. Overall diameter of 

 the holotype is 76.5 X 78.6 microns and the 

 spore body measures 53.1 X 55.2 microns. 

 There are two bladders between each radii 

 which are arranged so that the corners of 

 the spore body are not covered. The spore 

 body is levigate although it appears very 

 finely granulose under oil immersion. The 

 bladders are finely granulose. On the 

 proximal surface of the spore body there 

 are scattered several round papillae. The 

 trilete mark extends to the margin of the 

 spore wall and the lips and commissure are 

 not well developed. The spore coat is 1.5 

 to 2 microns thick and the bladders about 

 one micron thick. 



Holotype. — Maceration 519-A Slide 1, 

 Dekoven coal bed, Williamson County, 

 Illinois. 



Discussion. — In some forms it is difficult 

 to be certain whether there are five or six 

 bladders because one of the bladders may 

 not be clearly divided to the spore body. 

 Pollen grains from the modern genus Podo- 

 carpus upon occasion display unusual con- 

 ditions when a species which normally has 

 two bladders may possess three or four 

 bladders. This is thought to be due to the 

 failure of the mother cell to divide into four 

 pollen grains after the second division. In 

 some forms, one of the two bladders divides 

 resulting in three bladders ; in other forms 

 the two bladders divide resulting in four 

 bladders of equal proportions. Whichever 

 occurs, the body of the grain is larger than 

 that of normal pollen grains of the species 

 in question. This merely illustrates that 

 there may exist some variance in the normal 

 bladder number. 



Alati-sporites inflatus sp. nov. 



Plate 4, figure 2 



Description. — Spores are radial, body is 

 subtriangular in transverse plane, sub- 

 spherical in outline including bladder. 

 Margin of body wall between radii general- 

 ly is slightly convex or slightly concave and 

 the corners are broadly rounded. Overall 

 diameter of holotype is 120.4 X 129.6 mi- 

 crons and the known size range is from 



120 X 123 microns to 148.7 X 150 mi- 

 crons. The three bladders of the holotype 

 vary in the longest diameter from 87.1 mi- 

 crons to 93.4 microns. The bladders ex- 

 tend over the proximal side of the spore 

 body 8 to 15 microns and about the same 

 distance distally. The bladders are almost 

 devoid of folds and appear inflated. The 

 juncture of the bladder and the spore body 

 is somewhat irregular but in general follows 

 the outline of the spore body. The spore 

 coat is levigate with a few minor small 

 round thickened areas scattered on the 

 proximal surface. The bladders (peri- 

 sporial) are about 1 micron thick and 

 sharply granulose. The trilete mark ex- 

 tends nearly to the margin of the spore 

 wall, the lips and commissure are somewhat 

 developed. 



Holotype.— Maceration 543-C Slide 6, 

 No. 5 coal bed, Fulton County, Illinois. 



Alati-sporites punctatus sp. nov. 

 Plate 4, figure 4 



Description. — Spores are radial, spore 

 body is subtriangular in transverse plane, 

 overall transverse outline is irregularly 

 ovate to almost round. The margin 

 of the spore wall between radii is gener- 

 ally convex and the corners are some- 

 what bluntly pointed. The overall meas- 

 urement of the holotype is 102 X 98.75 

 microns while the spore body is 78.6 

 X 76.5 microns. The bladders vary in length 

 from 70 to 80 microns and from 25.5 to 29.7 

 microns in width. The bladders overlap 

 on the spore body about 5 microns. The 

 known size range of the spore body in the 

 mean diameter is from 70.5 to 79 microns. 

 The spore coat is obvermiculate (as ob- 

 served on undermacerated material) to 

 punctate (on overmacerated material) and 

 the bladders are finely granulose. Trilete 

 mark extends nine-tenths the distance to 

 the spore wall and the lips and commissure 

 are not developed. The spore coat is 4 to 

 5 microns thick and the bladders are 0.75 to 

 1.25 microns thick and greatly folded to 

 appear sometimes as though there were more 

 than three bladders. By focusing up and 

 down it is possible to prove the existence of 

 no more than three bladders. 



