40 



PENNSYLVANIAN SPORES OF ILLINOIS 



ly but widely scattered punctations. The 

 trilete rays are distinct and extend more 

 than three-fourths of the distance to the 

 margin of the spore wall. The lips are 

 slightly elevated and the commissure is 

 distinct. The spore coat is more than 2 

 microns in thickness. 



Holotype. — Maceration 587 Slide 13, 

 Battery Rock coal bed, Hardin County, 

 Illinois. 



Discussion. — T. priscus sp. nov. is the 

 first species of the genus recognized from 

 the Caseyville group in Illinois. 



Triquitrites protensus sp. nov. 

 Plate 8, figure 2 



Description. — Spores are radial, trilete, 

 and triangular in transverse plane. The 

 interradial margins are slightly concave or 

 convex and have arcuate thickenings. The 

 overall measurement of the holoytype is 

 37.8 X 36.5 microns, and the known size 

 range is from 33.5 to 39 microns. The 

 -arcuate thickenings appear to originate 

 from the proximal side of the spore equator 

 and their total width is 10.5 microns of 

 which 4.2 microns extend beyond the body 

 cavity. The length of the thickenings is 

 the same as the width, 10.5 microns. The 

 spore coat is levigate, but frequently small 

 fragments of debris are found clinging to 

 the proximal and distal surfaces. The 

 trilete rays are fairly distinct and the lips 

 are slightly elevated. The commissure may 

 or may not be clearly visible. The rays 

 extend almost to the margin of the spore 

 wall. The spore coat varies in thickness 

 from 2 to 3 microns. 



Holotype. — Maceration 519-B Slide 1, 

 Dekoven coal bed, Williamson County, 

 Illinois. 



Discussion. — The thickenings of T. pro- 

 tensus sp. nov. are distinct from those of all 

 other known species of the genus. 



GENUS Calamospora S. W. and B., 

 1944 



Plate 9, figures 1-5 



Calamospora is present in almost every 

 coal bed in Illinois. It is abundantly pres- 



ent in the coal beds of Tradewater, Car- 

 bondale, and McLeansboro age. Maxi- 

 mum abundance is recorded from the 

 Jamestown and Cutler coal beds of south- 

 ern Illinois. Calamospora is present in 

 moderate abundance in the Seville, Rock 

 Island, No. 2, No. 5, Shoal Creek, and 

 LaSalle coal beds. 



The genus is readily identified with one 

 exception : mildly ornamented species of 

 Punctati-sporites might possibly be confused 

 with Calamospora. 



Identification of species is not always 

 easy because of lack of ornamentation. 

 Also it is thought that the spores of the 

 genus are not readily segregated into re- 

 stricted generic relationships. Thus Cala- 

 mospora appears to be a somewhat general- 

 ized form, possibly representing several 

 types of parent plants. Hartung's (1933) 

 work suggested the Calamarian relation- 

 ship for which spores of this type have been 

 placed in the genus Calamospora. Arnold 

 (1945) described a fructification, Bow- 

 manites, which was heterosporous, and both 

 megaspores and microspores are of the 

 Calamospora type. Certainly Arnold's 

 small spores of this fructification (figs. 5 

 and 9) are referrable to Calamospora. 



Description of the genus Calamospora is 

 characterized as follows, based on eight 

 previous species and five new species: 

 Spores are radial, trilete, originally spheri- 

 cal in shape, and variously folded. The 

 known size range is from 30 to 165 mi- 

 crons. The ornamentation is in general 

 levigate. The trilete rays may extend from 

 one-fourth to more than two-thirds the 

 distance to the margin of the spore coat. 

 The lips may be greatly developed or lack- 

 ing, and the commissure may be thin or 

 wide. The area contagionis may be pres- 

 ent or absent. The spore coat is usually 

 very thin, 2 to 3 microns, and translucent. 

 Two species have spore coats up to 6 mi- 

 crons thick. Species with thin spore coats 

 usually are folded. 



It is possible that C. obesus (Loose) 

 comb. nov. S. W. and B,, 1944, and C. 

 flava sp. nov. are megaspores. The only 

 basis for this is their thick spore coats, 5 to 

 6 microns. However, this thickness is 



