35 



more closely spaced projections than P. 

 grandiverrucosus Kosanke, 1943, which it 

 otherwise resembles. 



Derivation of name. — The species name re- 

 fers to its coarse (rudis) ornamentation. 



Punctatisporites staplini sp. nov. 

 Plate 5, figures 5-6 



Diagnosis. — The spores are radial, trilete, 

 circular to subcircular in outline, and usual- 

 ly in good proximal-distal orientation. The 

 straight trilete rays, often obscured by orna- 

 mentation, are approximately equal to the 

 spore radius (as much as 31 /x long when 

 spore is obliquely compressed) . Lips are ab- 

 sent. The proximal and distal surfaces are 

 covered with coarse grana 1 to 2.5 /x in di- 

 ameter and about 2 /x high. Most of the 

 grana are rectangular in cross section and 

 have truncated flat ends, but some are cone 

 shaped. They are rectangular, circular, or 

 oval in end view. They are uniformly dis- 

 tributed, and there is not enough space be- 

 tween them for additional grana of equal 

 size. About 60 grana extend beyond the mar- 

 gin. The spore coat is about 1.5 /x thick. 

 Dimensions (26 specimens) : size range 42.1 

 to 57.7 fx in maximum diameter; median, 



51.4 ii (not including ornamentation). 



Holotype. — Plate 5, figure 5; negative 6116; 

 Fithian Gyclothem, maceration 1170-C, slide 

 16, coordinates, 126.1 X 37.7; size, 51.2 by 



50.5 /x (not including ornamentation) . 



Paratype. — Plate 5, figure 6; negative 5780; 

 Fithian Gyclothem, maceration 1170-G, slide 

 3, coordinates, 127.6 X 30.3; size, 51.8 by 

 48.9 /x. 



Comparison. — Punctatisporites staplini is 

 coarser grained than P. nahannensis Hacque- 

 bard & Barss, 1957, and the rays of P. nahan- 

 nensis are shorter, from one-half to two- 

 thirds of the spore radius (9.6 to 16 /x long) . 



Derivation of name. — The species is named 

 in honor of Dr. Frank L. Staplin, palynolo- 

 gist, Imperial Oil Company, Canada. 



Punctatisporites sp. 1 

 Plate 5, figure 7 



The spore is radial, trilete, and circular in 

 outline. The rays are distinct, straight, and 

 extend about two-thirds the length of the 



spore radius. The proximal and distal sur- 

 faces are covered with a fine reticulate pat- 

 tern in which the muri are 1 to 2 /x wide and 

 the lacunae about 6 /x in diameter. The exine 

 is about 1 /x thick. 



Figured specimen. — Negative 4731; Trivoli 

 Gyclothem, maceration 1128-G, slide 3, co- 

 ordinates, 122.8 X 44.7; size, 58.4 by 56.8 /x. 



Punctatisporites sp. 2 

 Plate 5, figure 8 



The spore is radial, trilete, and subcircular 

 in outline, and has two major folds parallel 

 to the margin. The rays are distinct, straight, 

 and at least 22 ^ long. Lips are absent. The 

 exine, about 3.5 /x thick, is levigate under 

 oil immersion objective. 



Figured specimen. — Negative 5570; Hen- 

 shaw Formation, maceration 1122-A, slide 

 11, coordinates, 138.8 X 40.2; size, 64.9 by 

 54.2 ii. 



Punctatisporites sp. 3 

 Plate 5, figure 9 



The spore is radial, trilete, and circular in 

 outline. The rays are distinct, straight, and 

 about 23 /x long. The proximal and distal 

 surfaces are punctate to vermiculate, and 

 when slightly out of focus appear reticulate. 

 Closer examination shows that the sculpture 

 consists of deep pits commonly connected by 

 shallow channels. The exine is about 2 /x 

 thick. 



Figured specimen. — Negative 6147; Fith- 

 ian Cyclothem, maceration 1170-C, slide 7, 

 coordinates, 126.5 X 45.5; size, 64.9 by 62.6 [i. 



Punctatisporites sp. 4 

 Plate 5, figure 10 



The spore is radial, trilete, and subcircular 

 in outline. The rays are distinct, straight, 

 about three-fourths the length of the spore 

 radius, and are bifurcated at their ends. Lips 

 are absent, but pyramidal areas are slightly 

 thickened. With oil immersion objective, the 

 slightly granular ornamentation of the spore 

 coat is discernible. The exine is about 4 /x 

 thick. 



Figured specimen. — Negative 6515; Hen- 

 shaw Formation, maceration 1122-A, slide 

 24 ZB, coordinates, 124.5 x 38.3; size, 75.3 

 by 69.7 }i. 



