ALATI-SPORITES 



33 



Laevigati-sporites deltoides (Ibrahim) 

 Ibrahim, 1933, Sporenformen des Aegir- 

 horizonts, p. 20, pi. 2, fig, 2. 

 Note. — The species is larger than most forms 

 assigned to Granulati-sporites. 



3. Granulati-sporites fistulosus (Ibrahim) 

 S. W. and B,, comb. nov. 



Reticulati-sporites fistulosus Ibrahim, 

 1933, idem, p. 36, pi. 5, fig. 35. 



4. Granulati-sporites gibbosus (Ibrahim) 

 S. W. and B., comb. nov. 



V errucosi-sporites gibbosus Ibrahim, 

 1933, idem, p. 25, pi. 6, fig. 49. 



5. Granulati-sporites granulatus Ibrahim, 



1933, idem., p. 22, pi. 6, fig. 51. 



Note. — Type species of Granulati-sporites. 



6. Granulati-sporites microgranifer Ibra- 

 him, 1933, idem, p. 22, pi. 5, fig. 32. 



7. Granulati-sporites microsaetosus 

 (Loose) S. W. and B,, comb. nov. 



Sporonites microsaetosus Loose, 1932, 

 Neues Jahrb., Beilage-Band 67, Abt. B, p. 

 450, pi. 18, fig. 40. 



Setosi-sporites microsaetosus (Loose) 

 Ibrahim, 1933, Sporenformen des Aegir- 

 horizonts, p. 26. 



Setosi-sporites microsaetosus (Loose) 

 Ibrahim, Loose, 1934, Inst. Palaobot. Arb., 

 vol. 4, no. 3, p. 148. 



8. Granulati-sporites microspinosus (Ibra- 

 him) S. W. and B., comb nov. 



Apiculati-sporites microspinosus Ibra- 

 him, 1933, Sporenformen des Aegirhori- 

 zonts, p. 24, pi. 6, fig. 52. 



9. Granulati-sporites parvus (Ibrahim) S. 

 W. and B., comb. nov. 



Sporonites parvus Ibrahim, 1932, Neues 

 Jahrb., Beilage-Band 67, Abt. B, p. 448, 

 pi. 15, fig. 21. 



Punctati - sporites parvus (Ibrahim) 

 Ibrahim, 1933, Sporenformen des Aegir- 

 horizonts, p. 21, pi. 2, fig. 21. 



Reticulati-sporites parvus (Ibrahim) 

 Loose, 1934, Inst. Palaobot. Arb., vol. 4, no. 



4, p. 154, pi. 7, fig. 13. 



10. Granulati - sporites piroformis Loose, 



1934, idem, vol. 4, no. 3, p. 147, pi. 7, fig. 19. 



11. Granulati-sporites (?) priddyi (Berry) 



5. W. and B., comb. nov. 



Zonales-sporites priddyi Berry, 1937, 

 Am. Midland Naturalist, vol. 18, no. 1, p. 

 156, fig. 2. 



12. Granulati-sporites torquifer (Loose) S. 

 W. and B., comb. nov. 



Sporonites torquifer Loose, 1932, Neues 

 Jahrb., Beilage-Band 67, Abt. B, p. 450, pi. 

 18, fig. 43. 



.Reticulati-sporites torquifer (Loose) 

 Loose, 1934, Inst. Palaobot. Arb., vol. 4, no. 

 - 3/.p. 154. 



13. Granulati-sporites trigonus (Ibrahim) 

 S. W. and B., comb. nov. 



Reticulati-sporites trigonus Ibrahim, 

 1933, Sporenformen des Aegirhorizonts, p. 

 ?>7, pi. 5, fig. 34. 



14. Granulati-sporites triquetris (Ibrahim) 

 S. W. and B., comb. nov. 



V errucosi-sporites triquetris Ibrahim, 

 1933, idem, p. 26, fig. 61. 



15. Granulati-sporites verrucosus (Wilson 

 and Coe) S. W. and B., comb. nov. 



Triquitrites verrucosus Wilson and Coe, 

 1940, Am. Midland NaturaHst, vol. 23, no. 

 1, p. 185, fig. 10. 



Genus Alati-sporites Ibrahim, 1933 

 Plate 1, figures 6-6b 



Symmetry. — Spores radial, trilete. 



Shape. — Body, exclusive of bladders, is 

 sub-triangular. 



Sise. — In the vicinity of 80 microns, 

 overall diameter. 



Ornamentation of spore. — Body moder- 

 ate ; may be punctate, finely reticulate ; 

 bladders smooth to punctate, etc. Blad- 

 ders, in general, less definitely ornamented 

 than spore body. 



Haptotypic structures. — Trilete rays 

 relatively long, little ornamented. 



Spore coat. — Consisting of two distinct 

 membranes; the outer (perisporal) mem- 

 brane is very thin and expanded in the in- 

 ter-radial areas to form bladders in a trim- 

 erous (in threes or multiples of three) 

 pattern. The spore body v^all (exospore) 

 is usually more than twice as thick as the 

 bladder membrane, darker, and easily 

 distinguished on that account. 



Affinity. — Spores of Alati-sporites seem 

 smaller but essentially similar in con- 

 struction to those of Spencerites (Scott, 

 1898, Kubart, 1910). The three inter- 

 radial bladders are most distinctive in 

 both genera. The significance of this 

 character is hardly well enough under- 

 stood to properly evaluate evidence as 

 to their relationship.^ 



^ The haptotypic relations, if any, of the three blad- 

 ders present on pollen of Podocarpus dacrydioides and on 

 Pherosphaera are not known, and comparison with Car- 

 boniferous forms does not seem particularly pertinent for 

 obvious reasons. 



It seems doubtful that the form described by Daugherty 

 (1941, p. 64) as Spencerites? chinleana is actually refer- 

 able to Spencerites or could be referred to Alati-sporites 

 since it apparently does not possess three interradial 

 bladders. Daugherty does not mention and the figure 

 given does not show the trilete sutures. If these were 

 present the species might be referred to Cirratriradites 

 provided the equatorial appendage was a flange. Lacking 

 these characteristics, his alternative suggestion of a 

 gymnospermic affinity seems most plausible. 



