72 CHELONTA. 



The inclusion of Manouria (Testudo emys), in which the caudal 

 shield is divided, in this genus l removes all grounds for the sepa- 

 ration of Colossochelys, the greatly produced epiplastral cornua of 

 the male of the latter not being of itself a generic character. 



Hadrianus, Cope 2 , from the Upper Eocene of North America, is 

 distinguished by the narrow vertebral shields ; the elongated neural 

 bones, which are hexagonal, with short postero-lateral surfaces 3 ; 



Fig. 12. 



Testudo ibera. — Carapace and plastron, reduced. The -j- + indicate the ex- 

 tremities of the axillary and inguinal buttresses. (From Boulenger's 

 ' Catalogue of Chelonians.') 



and the absence of alternation in the lengths of the extremities of 

 the costal bones, the caudal shield being divided. In some of the 

 undermentioned species from the Eocene and Miocene of Europe 

 the neurals and costals are unknown, and there is accordingly no 

 reason why they should not be referable to Hadrianus. 



1 See Boulenger, Catalogue of Chelonians, &c. p. 149 (1889). 



2 Proc. Amer. Phil. Soc. vol. xii. p. 468 (1873). 



3 See the figure of Hadrianus ( Testudo) corsoni by Leidy in Rep. U. S. Geol. 

 Surv. Terrs, vol. i. pt. i. pi. xi. (1873). 



