410 



Wirth and Stone: Diptera 



inner comb unarmed (fig. 14:2 It); pulvilli deeply bifid 



Polypedilum Kieffer 



— Base of 8th tergite and sternite of male subtruncate; 

 male dististyle with short bristles on the inner side, 

 some of which are microscopically forked near the 

 apex; fore tibia of both sexes with a round-ended 

 scale which has a minute apical spine or is unarmed; 

 usually both combs of hind tibia with a spine; pulvilli 

 entire Tanytarsus van der Wulp 



Larvae 



, Antennae not retractile; epipharynx present (fig. 

 14:27A); frontoclypeus narrow behind (fig. \A:S5d); 



labium 5 simple (figs. 14:27t, 14:35?) 2 



Antennae retractile; epipharynx absent; frontoclypeus 

 broadly rounded behind; labium complex (fig. 14:236) 



(PELOPIINAE) 6 



Tormae (premandibles) well developed (fig. 14:27A) 



3 



Tormae rudimentary (PODONOMINAE) 11 



, Paralabial plates usually absent (fig. 14:27i) or if 



present are not radially striated (fig. 14:25&,tf) 4 



Paralabial plates present and radially striated (figs. 

 14:31?, 14:32e) (TENDIPEDINAE) 25 



, Third antennal segment usually annulated (figs. 

 14:26t,Z; 14:29/), or if not, paralabial plates are present 



but not radially striated (DIAMESINAE) 13 



Third antennal segment not annulated (fig. 14:276, k, n); 



paralabial plates absent 5 



Restricted to intertidal rocks on seacoasts; preanal 

 papillae absent, instead 1-3 single hairs; anal gills 

 absent; basal bristle on labium absent; posterior pro- 

 legs short, only twice as long as broad (fig. 14:28Z) 



(CLUNIONINAE) 15 



Not restricted to intertidal rocks; preanal papillae 

 usually well developed and bearing more than 3 hairs; 

 anal gills and posterior prole gs usually well developed 



(fig. 14:27/) (HYDROBAENINAE) 18 



Body slender, segments with a few scattered bristles 

 (fig. 14:22;'); head about 0.45-0.66 times as broad as 

 long; labrum with numerous clavate bristles on anterior 

 margin (fig. 14:22a 1 ); labium without paralabial comb, 

 but with labial vesicles (fig. 14:22c); both pairs of 

 anal gills close to anal opening .. . Pentaneura Philippi 

 Body stout, segments with a longitudinal hair fringe on 

 each side; head broader, about 0.66-1.0 times as broad 

 as long; labrum with 6 sense vesicles on anterior 

 margin (fig. 14:23a); anal gills short, triangular, the 

 ventral pair remote from anus, on base or prolegs ... 7 

 Labium without paralabial comb, but instead with a 

 row of separate chitin points on each side (fig. 14:23c); 

 head about 1.5 times as long as broad; antennae 1/2 



to'3/4 as long as head 8 



Labium with paralabial comb (figs. 14:236, 1; 14:24d); 

 head short, never more than 1/3 longer than broad; 



antennae 1/4 to 1/3 as long as head 9 



Antennae very long, about 3/4 as long as head; man- 

 dibles hooklike Clinotanypiis Kieffer 



Antennae 1/2 as long as head; mandibles curved (fig. 



14:23p) Coelotanypus Kieffer 



Larva with 6 anal gills; lingua ("glossa") of hypo- 

 pharynx with 5 subequal yellow teeth ..Pelopia Meigen 



Larva with 4 anal gills 10 



Superlinguae ("paraglossae") of hypopharynx scalelike, 

 with several teeth on outer margin; lingua ("glossa") 



with 5 teeth (fig. 14:24/) Procladius Skuse 



Superlinguae of hypopharynx slender, with forked apex, 

 or if with toothed margin the lingua with 4 subequal 



yellow teeth (fig. 14:23c) Anatopynia Johannsen 



Labrum broad, the epistomal suture very weakly curved 

 cephalad; labral chaetae developed as short, fine 

 hairs (fig. 14:26d) 12 



H), 



11 



More correctly called the hypostome in recent morphological 

 papers. 



— Labrum narrow, the epistomal suture strongly curvec 

 cephalad; labral chaetae developed as 2 groups o 

 strongly curved, distally serrate bristles; 3 bristles 

 epipharyngeal comb Podonomus Philipp 



Boreochlus Edwards 



12. Tubercles bearing the labral bristles scarcely twice a< 

 high as broad (fig. 14:26d); preanal papillae as long ai 

 posterior prolegs, each with 13 bristles (fig. 14:26c) 

 head with 2 eyespots on each side; annulate part 

 antennae a little longer than the nonannulate 2nd seg- 

 ment (fig. 14 :26i) Lasiodiamesa Kieffei 



— Tubercles bearing the labral bristles about 5 times as 

 high as broad; preanal papillae much shorter than the 

 posterior prolegs, each with 6 bristles; head with 

 eyespots on each side; annulate part of antennae about 

 1/2 as long as the non-annulate 2nd segment (fig. 

 14:262) THchotanypus Kieffer 



13. Head capsule with warts or tubercles (fig. 14:29c, a 1 ); 

 abdominal segments with numerous small stellate 

 hairs (fig. 14:29o); posterior prolegs with sucker discs, 

 (fig. 14:29c) Heptagyia Philippi 



— Head and abdomen without such processes 14 



14. Paralabial plate with an arched margin, with or without 

 fringe of hairs, on each side of labium (fig. 14:256, d); 

 2 eyespots on each side of head. . . .Prodiamesa Kieffer 



— Paralabial plates absent Uiamesa Meigen 



Psilodiamesa Kieffer 

 Syndiamesa Kieffer 



15. Antennae 5-segmented (fig. 14:30/); tormae (preman- 

 dibles) with lateral teeth (fig. 14:30e); maxillae broader 

 than long, the palpi segmented; labial plate with a 

 lateral pair of subbasal setae (fig. 14:30A) 16 



— Antennae 4-segmented; tormae (premandibles) with 

 distal teeth of apex entire; maxillae not broader than 

 long, the palpi nonsegmented; labial plate without setae 



17 



16. Color of integument bluish or purplish; 2nd antennal 

 segment much shorter than the proximal segment; 

 labrum with a median sclerotized epipharyngeal bridge 

 Tethymyia Wirth 



— Color of integument greenish or whitish; 2nd antennal 

 segment about as long as proximal segment; labrum 

 without a median sclerotized epipharyngeal bridge 



Clunio Haliday 



17. Frontoclypeal suture absent; torma with broad flattened 

 lobe with an appressed hyaline frayed veil 



Thalassomya Schiner 



— Frontoclypeal suture present; torma with 3 blunt distal 

 teeth, without hyaline veil Telmatogeton Schiner 



Paraclunio Kieffer 



18. Antennae at least 1/2 as long as head, 2nd segment 

 slightly bent and darkened; posterior prolegs with a 

 basal ventral spur; 1st thoracic segment elongated 

 necklike, the 2nd and 3rd segments not separated (fig. 

 14:27e) Corynonewra Winnertz 



— Antennae not 1/2 as long as head, 2nd segment not 

 bent; spur on posterior prolegs absent; 1st thoracic 

 segment not elongated necklike (fig. 14:27/) 19 



19. Larva living in symbiotic relationship on body or under 

 wing covers of mayflies or stoneflies (fig. 14:31m) 

 20 



— Larva not symbiotic with mayflies or stoneflies 21 



20. Living under wing covers of mayflies; head unusually 

 small Symbiocladius Kieffer 



— Head not unusually small Hydrobaenus Fries (part) 



21. Bristle-bearing preanal papillae of last segment at 

 least twice as long as broad (fig. 14:27/) 22 



— Bristle-bearing preanal papillae shorter or entirely 

 absent (fig. 14:28?) 23 



22. Basal antennal segment slightly but distinctly bent 



Brillia Kieffer 



— Basal antennal segment straight (fig. 14:27£) 



Metriocnemus van der Wulp 



23. Labial plate with 11 teeth, the middle one broadly 

 truncated, hypopharynx with distally projecting hairs; 

 labrum with simple spines but without pectinate or 

 toothed plates; anterior prolegs fused, terminating in 



