426 



Wjrth and Stone: Diptera 



Fig. 14:38. Adult Heleidae. o-c, Culicoides cockerellii (Coq.): a, wing; b, mesonotum; c, male 

 genitalia, right gonostyles removed, d,e, Monohelea maculipennis (Coq.): d, wing; e, male geni- 

 talia. f,g, Dasyhelea cmcfo (Coq.): f, male genitalia; g, wing. h,i, Alluaudomyia-bella (Coq.): 

 h, wing; /, male genitalia. j,k, Alrichopogon occidental i s Wirth: ;", male genitalia; k, wing. I,m, 

 Forci'pomy/a macswaini Wirth: /, wing; m, male genitalia, n-q, Clinohelea usingeri Wirth: n, wing; 

 o, female hind tarsus, fourth and fifth segments; p, same, fore tarsus; q, male genitalia, right 

 gonostyles removed. r,s, Helea stigmalis (Coq.): r, wing; s, male genitalia, t, Dasyhelea grisea 

 (Coq.), larva; u, Alluaudomyia bella (Coq.), respiratory organ of pupa {a-s, Wirth, 1952; t,u. Thorn- 

 sen, 1937). 



14. Fifth tarsal segments armed below with straight, blunt, 

 black spines (batonnets) (fig. 14:39c) 15 



— Fifth tarsal segments without ventral batonnets (fig. 

 14:39rf), sometimes with sharp, tapering spines 18 



15. Claws of female unequal at least on mid and hind legs; 

 not California Dicrohelea Kieffer 



— Claws of female equal 16 



16. Femora armed below with short, stout, black spines 

 Sphaeromias Stephens 



— Femora unarmed 17 



17. Wing broad, especially at base, anal lobe expanded; 

 not California Jenkins helea Macfie 



— Wing narrow, anal lobe small. . J ohanns enomyia Malloch 



18. Femora armed below with strong spines; 5th tarsal 

 segments not inflated 20 



— Femora unarmed; 5th segment inflated on fore tarsi 

 (fig. 14:38p) 19 



19. Mid and hind legs of female with claws unequal or a 

 single claw with basal barb, 4.th tarsal segment deeply 

 bilobed and armed with spines (fig. 14:38o); costa not 

 produced beyond apex of 2nd anterior radial cell (fig. 



14:38n); pleura with transverse silvery band 



Clinohelea Kieffer 



— Claws equal on all legs; 4th tarsal segment cordiform; 

 costa produced past apex of 2nd anterior radial cell 



Neurohelea Kieffer 



20. Fore femora greatly swollen, the fore tibiae arched; 

 apical 3rd of hind tibiae swollen; hind tarsi of female 

 with a single, long basally barbed claw; wings fasciate; 



not California Heteromyia Say 



— Fore femora not swollen or, if moderately so, fore 

 tibiae not arched 21 



21. Femora and tibiae all armed with long, sharp scattered 

 spines; not California Echinohelea Macfie 



— Fore femora, and often mid and hind pairs, armed 

 ventrally with short, stout, black spines 



P alp omyia Meigen 



22. Fifth tarsal segments of female with long, blunt, black 

 spines (batonnets) below (fig. 14:39i); costa in female 

 extending nearly to tip of wing (fig. 14:39A); femora 

 unarmed Probezzia Kieffer 



— Fifth tarsal segments unarmed 23 



23. Cross vein r-m very short, basal sections of radius and 

 media very close together; M 2 strongly elbowed at base 

 in female; costa extending to wing tip; femora unarmed; 

 not California Stenoxenus Coquillett 



— Cross vein r-m longer, basal radial cell not unusually 

 narrow; costa to about 3/4 of wing length (fig. 14:39/) 

 Bezzia Kieffer 



Larvae 



1. All body segments with short spines; prothoracic pro- 

 legs present (figs. 14:39//; 14:41a) (FORCIPOMYUNAE) 



2 



— No spines on body segments; prothoracic prolegs 

 absent (fig. 14:38<) 3 



