SUBSECTION CAMAROPHYLLI 337 



Material Studied— California: Smith 3858, 8703, 9029, 9.564. Col- 

 orado: Shope 383. Oregon: Kauffman, Franklin, Sept. 19, 1915; Mnrrill 

 1009 (type, from Corvallis, Nov. 1911); Smith 27493, 27590, 27591, 

 27914, 28097. 



Observations — Smith has studied this species in great detail both 

 in favorable and unfavorable seasons along the Pacific Coast. It is usu- 

 ally found growing gregariously under spruce. It differs from the form 

 of H. pudorinus found in the eastern United States, apparently, in the 

 bright ochraceous base of the stipe, the pronounced tendency to stain 

 yellow and finally orange when bruised, and the pinkish-buff to 

 pinkish-cinnamon gills. Fruiting bodies in all stages of development 

 have been studied, and no indications of a gelatinous universal 

 veil were present. It is interesting to note that in many respects 

 this western variety corresponds to Fries' description of H. pudorinus 

 better than does the H. pudorinus of eastern United States or of most 

 European authors. Fries (1821) described the stipe as "glaber," ob- 

 viously referring to the lack of a veil. Later (1838) he stated "Proximus 

 H. eburneo, sed firmior, stipite sicco glabro et colore uniformi I. lute- 

 scenti-maculoso distinctus. Cortina nulla." In 1874 he stated of the stipe 

 "vellum nullum. Pileus non pelliculosus, interdum (ut caro alba) luteo- 

 maculatus." It is difficult to understand why a species with a distinct 

 gelatinous universal veil has been referred to H. pudorinus by certain 

 European investigators. The colors as Fries described them, the tend- 

 ency to stain yellow, and the stipe characters of both H. pudorinus Fr. 

 and H. fragrans Murr. are practically identical. The color of the gills in 

 the latter might serve to separate it. However, studying the situation at 

 every opportunity since our first publication ( 1939 ) , we have come to 

 the conclusion that H. fragrans can only be justified as a variety of the 

 Friesian species. 



204 



Hygrophorus pudorinus var. fragrans i. pallidus 



( Sm. & Hes. ) , comb. nov. 



Lloydia2: 38.1939 



Hygrophorus fragrans f. pallidus Sm. & Hes., Lloydia 2: 38. 1939. 



Pileus 5-9 cm broad, convex with an inrolled margin, at first "warm 

 buff" on the disc and nearly white on the margin, fading in age to 

 "pinkish buff" on the disc and becoming "cartridge buff" or more sordid 

 on the margin, gluten sometimes forming yellowish sordid patches over 

 the surface, surface viscid, glabrous or faintly fibrillose-streaked be- 

 neath the gluten, margin cottony tomentose. Context thick, white, 



