FLOWER OF PINGUECULA VULGARIS, ETC. 045 



the fact that the posterior middle line of the ovarian wall is a line of suture, 

 and in consequence that the small posterior lip of the stigma is potentially 

 a double organ.* In fig. 27, the posterior lip is normal, but the anterior 

 (large) lip is tripartite. Fig. 28 represents a leftf antero-lateral view of the 

 same pistil, showing a slightly marked lobule (Ibl) at the base of the antero- 

 median lobe ; the right antero-lateral fissure, however, was found to be uncom- 

 plicated by any such lobule. Fig. 29 exhibits a nearly anterior view of an ab- 

 normality very similar to the last ; but where a lobule occurs on either side of 

 the base of the middle anterior lobe, that on the left side (to the right hand in 

 the figure) being developed to about the same exent as the corresponding lobule 

 in the last abnormality, while the lobule on the right side is considerably more 

 distinct.]; I have represented in fig. 30 a pistil with the posterior lip somewhat 

 broader than usual, though undivided, and the anterior lip cleft down the left 

 side, thus exhibiting one antero-lateral fissure. § A very small notch is seen on 

 the right side, which possibly may be held as representing a right antero-lateral 

 fissure. In fig. 31 is seen an abnormality of quite another character. Here the 

 stigma is altogether undivided and almost quite regular, resembling a funnel the 

 walls of which are to a great extent turned inside out from reflection of the 

 margin. As I previously mentioned, when treating of the andrcecium, the two 

 staminodes here are well developed, with distinct filaments and anther-like 

 terminal knobs. 



Morphological Constitution of the Ovary. 



In connection with the monstrous pistils just described, and of course always 

 keeping in view the normal course of development, I would here make a few 

 observations as to the probable morphological constitution of the ovary. The 

 ordinary view has hitherto been, that the ovary in Lentibulariacese is bicarpellary, 

 a view supported by the bilabiate stigma, bivalved capsule, and last, not least, by 

 the fact that of the somewhat numerous vascular bundles entering its walls, the 

 two strongest are in the mesial plane, one anteriorly the other posteriorly. This 

 view, however, must be set aside in the face of developmental facts, which show 

 the posterior middle line to be a line of suture. If it be objected that the presence 

 of a strong vascular bundle in the posterior middle line constitutes a difficulty, I 

 need only point to the interpetiolar stipules in Cinchona, where we have a well- 

 marked vascular bundle occupying the middle line of the stipule, although that 

 middle line is the line of a suture, and not of a true midrib. There are, it seems 



* I have met with three instances of this bipartite condition of the posterior lip. 



f To the left of an observer supposed to stand in the axis of inflorescence. 



J I have in my possession a third example of an ovary with tripartite anterior lip, but as I 

 have been unwilling to remove the stamens from the specimen, I cannot say what appearance is pre- 

 sented on an anterior view; its posterior aspect, however, is almost identical with that given in 

 Plate XXX. fig. 27. 



§ This antero-lateral fissure is uncomplicated by any lobule. 



