



TROPHOLUM PEREGRINUM AND T. SPECIOSUM. 227 
or arrested punctum vegetationis, of the floral axis. A portion of the tube of 
the style (ts) is also shown, and this, it is to be noted, remains open, 7.¢., does 
not become filled up with “conducting tissue;” just as has been observed by 
ScHacut in 7. majus.* 
After fertilisation, I have observed the immediately succeeding stage where 
the germinal vesicle has undergone subdivision into two by a horizontal sep- 
tum, like what has been noticed by ScuAcut in 7. majus. 
The stages immediately following this I have not seen, and the next observed 
is that represented in Plate XIV. figs. 2 and 3. Here the germ (g) is multicel- 
lular and pyriform with pointed base (0) fitting into the apex of the embryo-sac. 
On side view this pyriform germ exhibits about 50 cells. At this stage is very 
distinctly to be noted a single layer of nucleated particles, pretty uniformly dis- 
tributed and embedded in the protoplasm which lines the inner surface of the 
embryo-sac. This layer, which I have attempted to indicate in Plate XIV. 
fig. 3 (ed), is interesting as undoubtedly representing a transitory endosperm 
which, as is known, is almost, if not altogether absent in 7. majus. 
In Plate XV. fig. 12, the germ is represented in the immediately succeeding 
stage, where it assumes the flask or soda-water bottle form, almost exactly like 
that observed in 7. majus by HorMEISTER, SCHACHT, and myself, the only import- 
ant difference being in its consisting of a much larger number of cells than that 
in 7. majus,—this standing in relation to the stouter and more massive general 
character of the germ of 7. speciosum, as compared with that of the common 
species. In 7. majus the flask form is assumed at a very early period, as may 
be seen in the figures after HormeIsTEer and Scuacut (Plate XV. fig. 18, A, B and 
C); that at A giving a side view of only half a dozen cells. 
In the next stage (Plate XIV. fig. 4 and Plate XV. fig. 13) the flask-shapea 
germ has become somewhat elongated, particularly in the neck and pointed 
base, in which parts the cells are becoming slightly enlarged. As yet the axis 
of the germ is straight. The chalazal extremity of the embryo-sac (Plate XIV. 
fig. 4) now exhibits signs of that enlargement which goes on progressively until 
maturation of the fruit, the embryo proper with its large amygdaloid coty- 
ledons being lodged in this chalazal dilatation of the sac. Otherwise, the 
young seed at this stage hardly differs from that represented in Plate XIV. fig. 
2, except in its somewhat greater size. 
In Plate XIV. fig. 5, the micropylar extremity of the young seed and 
embryo-sac are seen to have become much more markedly curved, and coinci- 
dently with this (perhaps in consequence of it) the now still more elongated 
germ exhibits a distinct knee-like bend above its middle. The elongation of the 
neck and pointed base of the germ by enlargement of their constituent cells is 
now more distinctly marked, these regions appearing transparent by contrast 
* Scnacut, Bot. Zeit. 1855, pp. 641-2. 
VOL, XXVII. PART II. oN 
