A REVISION OF THE ETHIOPIAN DREPANIDAE (LEPIDOPTERA) 41 



Ground-colour of surface of fore wing (Plate 4, fig. 283) yellowish grey (holotype) or grey, 

 lustrous ; dark markings yellowish brown (holotype) or dark reddish brown, non-lustrous ; 

 conspicuous black marking at anal margin (absent in lectotype and paralectotype ; also present 

 in holotype of heringi). Ground-colour of hind wing similar to fore wing but heavily speckled 

 with dark brown and buff (holotype) or dark brown and pink ; postmedial fascia pale buff ; 

 subterminal markings dark brown, non-lustrous. 



Costal area under surface of fore wing greyish buff ; rest of wing (except for pale buff area 

 overlapped by hind wing) buff (holotype) or pink, becoming more greyish and striate with dark 

 grey apicad and distal to postmedial fascia ; postmedial fascia and discocellular spot dark grey 

 (holotype) or black. Under surface of hind wing pale buff with grey outer marginal area, pink 

 between grey area and cell in male from Mabira Forest ; discocellular spot as for fore wing ; 

 postmedial fascia grey, weakly marked, strongly convex distally. 



Abdomen grey dorsally, buff ventrally. 



cJ genitalia (Text-figs. 58-60) : valve process long, stout, arcuate ; posterior processes of 

 uncus truncate. 



$ (Plate 4, fig. 284) . Similar to male but outer marginal process absent on fore wing and much 

 reduced on hind wing, and with postmedial fascia and subterminal fascia on upper surface of 

 hind wing differently shaped. (Ground-colour of upper surface of wings of specimen from 

 Entebbe reddish buff, and with dark marking at middle of anal margin on fore wing.) 



$ genitalia (Text-fig. 61) : dorsal lobes of eighth segment not striate medially ; ninth seg- 

 ment well sclerotized laterally and ventrolaterally. 



Measurements. A.P.R. : <J, $. IQ - Wing : <$. 20-0 mm. (2) ; ?. 22-5, 220-23-0 mm. 

 (2). 



Discussion. This species is closely related to the West African heringi. The 

 colour-pattern of the wings is probably very similar to that of heringi although the 

 latter is unfortunately known only from the worn holotype. The male genitalia, 

 however, readily separate these two species : the shape of the valve processes, uncus 

 and eighth sternum are particularly diagnostic. Until more material of both sexes 

 is available I consider it better to treat heringi and heterogyna as species of a super- 

 species than as subspecies of a single species. 



A recent paper by Stempffer and Jackson (1962) has shown that the aquatic 

 barrier between Bugalla Island (the type locality of heterogyna) and the mainland of 

 Africa is sufficient to bring about subspeciation in three butterfly species. It is 

 interesting in this respect to find no evidence of significant geographical variation in 

 the few available specimens of heterogyna. The slight variation in coloration 

 mentioned in the description may be due to fading of the original Hampson type 

 material from Bugalla. 



Material examined. Type. I have selected and labelled the male syntype as 

 the LECTOTYPE. It bears the following data " Uganda, Victoria Nyanza, Sesse 

 Is., Bugalla I., Carpenter, 1914-276 ; Metadrepana heterogyna Hampson type <J 

 Hmpsn ; Drepanidae genitalia slide No. 1153 " ; in the British Museum (Natural 

 History) . 



Paralectotype. 1 $, with same collection data as lectotype. 



Other material. Coryndon Museum, Nairobi. Uganda : 1 $, Jinja, Mabira 

 Forest, x.1962 (Carcasson) ; 1 $, Entebbe, ix.1964 (Burgess). 



