A REVISION OF THE ETHIOPIAN DREPANIDAE (LEPIDOPTERA) 115 



$ genitalia : In boboa (species-group fenestrata) the bursa is ornamented with an irregular 

 band of scobinations ; in unidentified females of species-group spicata the bursa bears a T-shaped 

 signum. 



Discussion. Crocinis was erected for three new species : fenestrata (selected as 

 type-species by Butler) plana, and ochracea. [The latter two species have been 

 transferred to the family Geometridae by previous authors.] Drapena was 

 established by Gaede (1927 b : 189) for Drapena forata Warren (selected as type- 

 species by Gaede) and Drepana tetrathyra Mabille. Examination of the holotypes of 

 fenestrata and forata has shown them to be conspeciftc and that Drapena, 1927, must 

 be treated as a junior synonym of Crocinis, 1879. In this revision eight new 

 Madagascan species are added to the genus Crocinis, and tetrathyra Mabille is trans- 

 ferred to this genus from Gogana Walker (an Indo-australian genus which bears 

 little resemblance to Crocinis). 



This Madagascan genus is probably most closely related to the African Spidia 

 Butler, from which it may have been derived, but can be separated from it in the 

 male genitalia by differences in the form of the valves (or vincular processes) and 

 saccus, the absence of a gnathus, and the presence of a well-developed anellus and 

 an only slightly modified eighth abdominal sternum. The female signum is probably 

 also diagnostic, but the females of only one species of Crocinis have been definitely 

 identified. 



The species spicata and licina are apparently identical externally but are separated 

 by small differences in parts of the male genitalia. They are sympatric, however, 

 in the forest of Anamalazoatra, near PeVinet, and are therefore probably distinct 

 species, not subspecies of one species. 



Considerable individual variation in coloration and in the size and shape of the 

 hyaline patches is present in spicata and viettei. Similar variation may exist in the 

 remaining species of this genus but there is insufficient material to show this. 



Several females of the species-group spicata in the Museum national d'Histoire 

 naturelle, Paris, the Institut scientifique, Madagascar, and the British Museum 

 (Natural History) can not, at present, be associated with the corresponding males. 

 One of these females is illustrated (Plate 13, fig. 325). 



The species of Crocinis can be readily separated into two groups which are diag- 

 nosed below. I have termed them species-group fenestrata and species-group spicata. 



Species-group fenestrata 



Outer margin of hind wing concave posteriorly (except felina) ; cell-spots on both surfaces of 

 fore and hind wing absent or weakly marked, q* genitalia : posterior digitate process of valve 

 (or derivative of vinculum) without sclerotized spine ; diaphragma without invagination ; uncus 

 simple, not bilobed at base ; vesica of aedeagus with cornutus or cornuti. 



Species-group spicata 



Outer margin of hind wing straight or slightly convex posteriorly ; cell-spots usually well 

 marked, <J genitalia : posterior digitate process of valve with sclerotized spine ; diaphragma 

 with anteriorly directed invagination medially ; uncus hood-like, bilobed at base ; vesica 

 scobinate but without cornuti. 



