DIPTERA FROM NEPAL 



CRYPTOCHAETIDAE, DIASTATIDAE 

 & DROSOPHILIDAE 



ByT. OKADA 



CONTENTS 



Synopsis ...... 



Introduction ..... 



Abbreviations used in the Text-figures 

 Acknowledgements .... 



Family Cryptochaetidae 



Family Diastatidae .... 



Family Drosophilidae 



Subfamily Steganinae 

 Subfamily Drosophilinae . 

 Geographical Distribution 

 Ecological Distribution 

 Notes on some Special Systematic Characters 



A. Archestinic characters .... 



B. Homeotic expression of the hooked scaly bristles 



C. Homology in the components of phallic organs and the systematic 



analysis of phallic formulae 

 References ...... 



Index ....... 



Page 



3 



4 



5 



6 



6 



8 



14 



15 



37 



112 



117 

 119 

 119 



120 



121 

 I2 4 

 I27 



SYNOPSIS 



The Cryptochaetidae, Diastatidae and Drosophilidae collected by Mr. Ralph L. Coe, British 

 Museum (Natural History), during the British Museum East Nepal Expedition, 1961-1962 

 amounted to twelve hundred and eighty-seven well-preserved dried flies and a puparial case. 

 They represent seventy-six species belonging to fifteen genera, including Lissodrosophila gen. n. 

 and thirty-seven new species. In addition to describing new forms, insufficiently known species 

 have been re-described according to current taxonomy. Brief notes of the geographical and 

 ecological distributions of the genera and species concerned are given, and some systematically 

 important problems concerning the occurrence of " archestinic " characters in a taxon, the 

 homeotic and compensatory expression of characters, and homology in the components of phallic 

 organs are analysed and discussed. 



