122 H. STEMPFFER 



Male genitalia (Text-fig. 108, lateral view ; Text-fig. 109, ventral view). Uncus composed 

 of two long digitate processes with rounded apices ; no subunci ; tegumen deeply notched on 

 its posterior margin ; vinculum fairly broad ; valves oblong, the upper edges almost straight, 

 the lower ones convex ; lower fultura composed of two small divergent arms fused basally to 

 the base of the valves ; in addition, there is an offshoot, which I consider an upper fultura, 

 articulated at about half-way along the upper edge of the valves, where it is slightly notched, 

 connected with the edge of the tegumen and entirely surrounding the penis ; penis verv 

 elongate, swollen at the base, the external portion tapering regularly and slightly curving ; 

 there are a few fine, short hairs on the uncus ; the valves are densely hairy, especially on the 

 lower edges and apices. 



The male genitalia of D. demeter Clench are identical with those of D. hymen. 



List of Species of Dapidodigma 



*Dapidodigma demeter demeter Clench, 1961, Ann. Carneg. Mus. 36 : 64, figs. 

 Dapidodigma demeter nuptus Clench, 1961, I.e. : 65, fig. 

 *Dapidodigma hymen (Fabricius), 1715. Fig. Cramer, 1779, (as liger). 



Genus IOLA US Hiibner 



When I undertook the revision of the " Iolaus complex ", in collaboration with 

 Neville Bennett, I found myself faced with considerable difficulties. The group is 

 so remarkably uniform in external appearance that its species are recognisable at a 

 glance ; the upperside is nearly always blue, the underside pure white with a pattern 

 of markings reduced to delicate postdiscal and antemarginal lines, and the hind wing 

 carries delicate tails at the ends of veins 2 and ib. But in respect of venation and 

 male genitalia this uniformity is entirely lacking. 



The genera hitherto widely employed within the Iolaus group have been based in 

 the main on the venation, and can be grouped as follows : — 



A. Fore wing with 10 veins in both sexes : — 



Hemiolaus, Stugeta, Pseudiolaus. 



B. Fore wing with 11 veins in both sexes : — 



Tanuelheira, Aphniolaus, Iolaus (sensu stricto), Epamera, Sukidion. 



C. Fore wing with 12 veins in the male, 11 in female : — 



Trichiolaus, Argiolaus. 

 Some of these genera seem to me to have been founded on characters of secondary 

 importance, for example, Trichiolaus on the hairiness of the eyes, Tanuetheira on the 

 great length of the tail, Aphniolaus on its yellow coloration ; and it is impossible 

 on any sound criterion to separate Iolaus (sensu stricto) from Epamera. 



However, examination of the morphology of the male genitalia demonstrates that 

 the great majority of species forming the Iolaus group have an armature of Thecline 

 type, that is to say, with the dorsal elements forming a kind of hood with well 

 developed subunci. Yet some others (e.g. Argiolaus) display a very different 

 structure : in the silas group, the uncus is divided into two long sharp points, the 

 subunci are rudimentary and the tegumen reduced ; parasilanus has a pseudotergum 



