GENERA OF AFRICAN LYCAENIDAE 151 



Genus APHNAEUS Hubner 



Aphnaeus Hubner, 1826, Verz. bek. Schmett. : 81 ; Aurivillius, 1898 : 327 ; 1924 : 407 ; 

 Pinhey, 1949 : 104. Type-species : Papilio orcas Drury, 1782, designated by Scudder (1875, 

 Proc. Am. Acad. Arts Sci. 10 : 116). 



Eyes clothed in dense short hair ; palpi fairly long, parallel, second segment long, slightly 

 ascending, clothed with dense scales, third segment very short, slender, horizontal, with blunt 

 apex ; antennae three-fifths the length of the costa, club progressively swollen, not well 

 differentiated ; q" Iore leg with tibia as long as femur, tarsus unsegmented, femur bearing long 

 black hairs, tibia and tarsus bearing strong spines ; mid and hind legs, femora bearing long 

 black hairs, tibiae as long as femora, metatarsi very long. 



Wing shape. Fore wing subtriangular with pointed apex ; hind wing much produced at 

 the anal angle, a delicate tail at the end of vein 2, a longer tail at the end of vein ib, an anal 

 lobe, the abdominal margin excised between the lobe and the end of ia. 



Wing venation (Text-fig. 299). Fore wing with 12 veins. 



Male genitalia (Text-fig. 135 : side view ; Text-fig. 136 : dissected, with the parts separated 

 and spread out ; Text-fig. 137 : view of the valves detached from the vinculum and rotated 

 through 180 in order to give a better view of the median band). Uncus reduced to a narrow 

 strip bordering the edge of the tegumen ; subunci long, bent in an acute angle, apices blunt ; 

 tegumen subrectangular, hood-shaped ; vinculum fairly broad with a short rounded saccus ; 

 lower fultura composed of two flattened arms attached to the valves about midway, not to their 

 base near the vinculum ; valves elongate, the upper and lower processes distinctly separate in 

 their distal half and both having rounded apices, the upper process being more slender and a 

 little longer than the lower one ; a weakly sclerotized band unites the two upper processes, 

 passing above the external portion of the penis, which thus lies between the body of the valves 

 and this band ; penis large, with a protuberance on its upper surface situated a little before the 

 wall of the genital cavity, the external portion tapering gradually to a blunt apex ; the vesica 

 enclosing in its base a tuft of long, strong spines, and midway and at the apex a group of 

 cornuti ; uncus and distal portions of the valves pilose. The genitalia of the other species of 

 Aphnaeus are similar to those of orcas. 



This peculiar arrangement of valves sheathing the penis is not confined to the 

 genus Aphnaeus, similar structures being found in the allied genera Cigaritis (palae- 

 arctic), Apharitis, Spindasis, Chloroselas, Zeritis, Desmolycaena, Axiocerses, Phasis 

 and Erikssonia, and in nearly all species of these genera the markings on the under- 

 side of the wings show some likeness to those of species of Aphnaeus, e.g. the presence 

 of metallic or nacreous bands, spots or dots. The median band uniting the valves 

 may easily escape observation because it is very little sclerotized, translucid, and so 

 fragile that it is difficult not to break it when dissecting out the penis ; when broken 

 it is easy to mistake it for some membranous fragment. 



List of Species of Aphnaeus 

 * Aphnaeus adamsi Stempffer, 1954 : 513, fig. and fig. genitalia. 



* Aphnaeus affinis ajfinis Riley, 1921, Trans, ent. Soc. Lond. 54 : 249, figs. Fig. 



genitalia, Stempffer, 1954. 



* Aphnaeus affinis seydeli Berger, 1952, Lambillionea 52 : 68. Fig. Stempffer, 



1954- 

 *Aphnaeus argyrocyclus Holland, 1890. Fig. Holland, 1893. Fig. genitalia, 

 Stempffer, 1954. 

 propinquus Holland, 1893. 



