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H. STEMPFFER 



cavity ; subunci rather short, almost straight, tapering gradually from their base to the blunt 

 apices ; tegumen reduced to a rather narrow median strip ; vinculum wide above, narrow 

 below ; lower fultura consisting of two slender curved arms arising from the base of the valves ; 

 valves oblong, proximal portion oval, apex serrated, the size of the teeth increasing gradually 

 from the upper to the lower process, the lower edge deeply excised near the apex ; penis of 

 the same structure as in the genus Syntavucus, i.e. short, massive with carina penis and cuneus, 

 the latter bifid ; uncus and lower processes of valves pilose. 



As can be seen by comparing figures, the male genitalia of C. webbianus are closely 

 analogous to those of Ethiopian Syntarucus and American Leptotes. 



In his generic description, Butler included in Cyclyrius the species aeqaatorialis 

 Sharpe and juno Butler ; Aurivillius (1924 : 471-473) also included in what he 

 called the Cyclyrius group, tsomo Trimen, noquasa Trimen, kisaba Joicey & Talbot, 

 notobia Trimen and crawshayi Butler. Later on other species or subspecies were 

 placed in this genus by other authors, viz : boma Bethune Baker, ruandensis Joicey 

 & Talbot, vidcanica Joicey & Talbot, marungensis Joicey & Talbot, wollastoni 

 Bethune Baker and major Joicey & Talbot. 



I have examined the male genitalia of all these species or subspecies. As I have 

 shown (1943, Annls Soc. ent. Fr. 1942 : 125) they have nothing in common with 

 those of C. webbianus ; crawshayi is a Euchrysops (Stempffer, 1938, Mission Omo 

 4: 204, fig. 26) and all the others belong to the genus Harpendyreus. They must be 

 excluded from the genus Cyclyrius, which they resemble only fortuitously and in the 

 markings on the underside of the hind wing. 



On the other hand mandersi H. H. Druce, from Mauritius, described as a Nacaduba, 

 is a true Cyclyrius not only in its venation but also in its male genitalia, of which I 

 give a drawing (Text-fig. 191) : it is true that the subunci are finely spinose, but the 

 penis has the highly characteristic form of Cyclyrius, Syntarucus and Leptotes. It 

 seems impossible to have any doubts as to the close relationship of these three genera 



Fig. 190. Cyclyrius webbianus (Brulle), <J genitalia. 



