96 R. W. CROSSKEY 



stage as well as the male adults, and many supposed species or subspecies of doubtful 

 validity have been described that are based upon small differences in pupal gill form 

 or the structures of the male genitalia. 



Rubzov (1959-1964 : 392-416) recognizes three species-groups (equinum-gvoup, 

 salopiense-gvoup, and sergenti-group) in Wilhemia, but it is here considered that there 

 is such a homogeneity among the constituent elements in the subgenus (genus sensu 

 Rubzov) , whatever their taxonomic ranking in status, that no species-group classifi- 

 cation is required ; I therefore recognize no species-groups in the present work. 



Enderlein (1921, 1922, 1925, 1930) assigned several non-Palaearctic species to 

 Wilhelmia but later removed some of them to other genera. None of these belong 

 in the subgenus Wilhelmia as now understood, but for convenient reference the 

 species involved are given in the following list with the subgenus to which they now 

 belong indicated in brackets after the specific name : exiguum Roubaud (Notolepria 

 Enderlein) ; hippovorum Malloch (Hemicnetha Enderlein) ; mallochi Enderlein (? 

 Simulium Latreille s. str.) ; pictipes Hagen (Hagenomyia Shewell) ; pygmaea 

 Enderlein {M or ops Enderlein) ; vittatum Zetterstedt (Psilozia Enderlein) ; wolff- 

 huegeli Enderlein (? Psilopelmia Enderlein). Enderlein (1930 : 93) placed australi- 

 ensis Schiner in Wilhelmia, but this a New Zealand species now belonging in the 

 genus Austro simulium Tonnoir. 



Included taxa. The subgenus includes all species and subspecies placed in 

 Wilhelmia in Rubzov's (1959-1964) monograph of Palaearctic Simuliidae, except 

 for Simulium guimari Becker (which, following the lectotype designation made for 

 this species by Crosskey (19656 : 669), belongs in the subgenus Eusimulium). In 

 addition it includes Simulium {Wilhelmia) zetlandense Davies, 1966, and Simulium 

 (Wilhelmia) balcanicum ssp. severinense (Dinulescu, 1966) which have been described 

 since the completion of Rubzov's monograph. 



The included taxa in the purview of the present paper are from North Africa and 

 the Canary Islands and are : Simulium (Wilhelmia) ariasi Seguy ; S.(W.) bar- 

 baricum Seguy ; S.(W.) canariense Seguy ; S.(W.) equinum (Linnaeus) ; S.(W.) 

 mediterraneum Puri ; S.(W.) pseudequinum Seguy ; S.(W.) quadrifila Grenier, 

 Faure & Laurent ; S.(W.) sergenti Edwards. [Note : it is almost certain that the 

 names of some of these taxa recorded or described from the North African area are 

 synonyms of each other.] 



Subgenus METOMPHALUS Enderlein 



Metomphalus Enderlein, 1935 : 361. Type-species : Metomphalus caffer Enderlein, 1935 

 [=Simulium medusaeformis Pomeroy, 1920], by original designation. 



Diagnosis. 5*? : Basal section of radius haired. Pleural membrane bare (some except- 

 ions). Katepisternum bare. Fore tarsus not or slightly dilated, fore basitarsus five to seven 

 times as long as its greatest breadth. Hind basitarsus 1 -4-1 -6 times as long as remainder of hind 

 tarsus (cf. Wilhelmia). $ : cibarium unarmed or occasionally with small denticles between 

 cornuae. Tarsal claws without basal tooth. Scutum with inconspicuous dark pattern, usually 

 in form of lyrate mark. Abdomen evenly covered with scales. Seventh sternite undeveloped. 

 Gonapophyses slender and pointed, usually curled but sometimes long and straight. Paraprocts 



