SIMULIIDAE OF AFRICA 107 



tarsi slender and the base of the radius bare. Tetisimulium resembles Hagenomyia 

 in having a well developed and rather hairy seventh sternite in the female and in the 

 elongate styles of the male hypopygium, in the shape of the larval postgenal cleft, 

 and to a lesser extent in the form of the cocoon and pupal gill (although in 

 Hagenomyia gill there are nine filaments and not six or eight), but it differs by having 

 the cibarium toothed (unarmed in Hagenomyia) and in the form of the larval mandi- 

 bular comb-teeth (all very large in Hagenomyia) ; it resembles Psilozia in having 

 the cibarium toothed, in the simple larval rectal gills and in the shape of the female 

 gonapophyses (slightly produced but rounded) , but differs from Psilozia in the long 

 styles with single spinule (in Psilozia the male styles are short and subquadrate with 

 2-4 apical spinules), in the larger pointed larval postgenal cleft, in the differently 

 formed larval mandibular comb-teeth and in cocoon and pupal gill form. 



The Palaearctic subgenus Obnchovia Rubzov has an almost identical distribution 

 to that of Tetisimulium, being largely confined to the more arid parts of the region 

 and particularly characteristic of the black-fly fauna of Soviet Central Asia : but 

 Obuchovia is not known from North Africa, and probably does not occur there. 

 Obuchovia has a superficial resemblance to Tetisimulium because of its haired pleural 

 membrane, bare base to the radius, often slender fore tarsi, almost simple female 

 claws, patterned female abdomen, and elongate heavy male styles, but there is 

 almost certainly little if any close relationship : in Obuchovia the male ventral plate 

 is flat and rounded, the pupal gill very short and concealed within the closely woven 

 long-necked cocoon, there are no dorsal spine-combs on any of the terminal pupal 

 abdominal segments, the form of the larval abdomen, mandibular comb-teeth and 

 hypostomium are quite different, the female gonapophyses are elongate tongue-like 

 and the cibarium is unarmed. 



The species of Tetisimulium form a homogeneous subgenus, and no species-groups 

 are recognized. 



Included taxa. All taxa placed in Tetisimulium by Rubzov (1959-1964), of which 

 only one species occurs in North Africa (Morocco and Algeria) : Simulium {Teti- 

 simulium) bezzii (Corti). 



[Note : the publication date for Melusina bezzii Corti has always been cited 

 erroneously in the taxonomic literature as 19 16, but the name bezzii is nomen- 

 claturally available from its earlier citation in the key given by Corti (1914 : 197, 

 198), and the correct date for the species is 1914. Simulium atlas Seguy, described 

 from Morocco by Seguy (1930), is a synonym of bezzii : synonymy established by 

 Grenier & Theodorides (1953 : 439)]. 



Subgenus ODAGMIA Enderlein 



Odagmia Enderlein, 1921 : 199. Type-species : Simulia ornata Meigen, 181 8, by original desig- 

 nation. 



Discosphyria Enderlein, 1922 : 72. Type-species : Discosphyria odagmiina Enderlein, 1922, by 

 original designation. 



Diagnosis. c?9 : Basal section of radius bare. Pleural membrane haired. Katepisternum 



