34 R- W. CROSSKEY 



Australian fauna, and two other subgenera may be represented by species here 

 left subgenerically unplaced (these are hippobosca Paramonov whose characters 

 are very close to those of the Palaearctic subgenus Phorantha Rondani, and 

 nigrisquama Malloch that has characters similar to those of subgenus Brumptallophora 

 Dupuis) . Several of the Australian species, however, do not fit any of the Palaearctic 

 segregates, but instead are assignable to Mormonomyia Brauer & Bergenstamm, 

 a subgenus of Alophora s.l. until now considered to be entirely African. A careful 

 comparison of these Australian species with the African species (including the 

 type-species of Mormonomyia) has shown a complete concordance in the essential 

 characters, and it is here considered that the Australian species must be placed as 

 consubgeneric with the African ones; the subgeneric name is therefore applicable 

 to these Australian species. (Here it may usefully be noted that comparison of 

 the New Zealand genus Campbellia Miller with Mormonomyia, made whilst comparing 

 Australian and African species, has shown no differences that can be considered 

 of any nomenclatorial consequence and the name Campbellia ought almost certainly 

 to be placed in synonymy with Alophora [Mormonomyia).) 



In addition to Alophora s.l. the Australian Phasiini includes the genera Efftayloria 

 and Besserioides. The former occurs also in New Guinea and is closely allied to 

 Alophorophasia Townsend (synonym: Kosempomyia Villeneuve), having the lower 

 surfaces of the femora similarly armed with rows of short stubby spinules. 

 Besserioides is entirely Australian on present evidence, but is evidently very closely 

 allied to the monotypic Ethiopian genus Bogosiella Villeneuve (from which it differs 

 by having the frons equally wide in both sexes and by having the posterior spiracles 

 of the puparium on prolonged paired bosses) . 



The main external features of Phasiini are as follows. Head holoptic or dichoptic, some 

 eye facets of both sexes enlarged in some forms; § without proclinate orbital setae; face 

 without a median keel; ocellar setae proclinate or absent; antennae usually short or very 

 short, if moderately long (e.g. Alophorophasia alata Townsend) nevertheless falling well short 

 of epistomal margin; palpi present, sometimes weak; humeral callus with 0—2 setae; acr setae 

 usually o + 1 (hair-like); dc setae variable (often hair-like), commonly 1 + 2 (sometimes 3) 

 post dc or only one distinct; one sa seta; one post ia seta or none; usually 1 + 1 or o + 1 stpl 

 setae, rarely 2 -f- 1 ; inf rasquamal hairs present or absent ; scutellum with one (apical) or 

 two (basal and apical) pairs of marginal setae, rarely weak third pair, without discal setae; 

 scutellum sometimes rather bullate and postscutellum often much produced posteriorly and 

 then visible beyond the scutellum when seen from above; wings either of normal form or 

 (very commonly) much produced in the anal region (especially in £) so that they are very 

 broad basally and short; wings often wholly or partially infuscate or yellowish or orange 

 basally ; second costal sector bare ventrally (first sector often also bare as well) ; basal node 

 of i? 4 , 5 usually with one or two very minute hairs, sometimes bare ; bend of M forming an 

 even curve without trace of appendix; cell R 5 usually closed well before the wing margin 

 and petiolate, occasionally just open or closed at the margin (closed and petiolate in known 

 Australian forms) ; last section of Cwj very short, less than half as long as m-cu (sometimes 

 postero-apical corner of discal cell virtually reaching the margin of the wing) ; lower calypter 

 very broad posteriorly, hind margin nearly straight but outer posterior corner sometimes 

 strongly produced; legs weakly bristled in most forms or devoid of definite setae, sometimes 

 with a few stubby setae or combs of stubby setulae ventrally on the femora; tibiae often 

 strongly curved or appearing so, hind tibia without scale fringes; hind tibia without pd 

 preapical seta or pv apical seta; hind coxae close to or at least not very remote from the 



