88 R. W. CROSSKEY 



- Inner vertical setae subparallel. Abdominal Ti + 2 and T3 each with one pair of 



median marginal setae (those of Ti + 2 sometimes hair-like or virtually absent). 

 Abdomen usually with one or two distinct discal setae on one or both intermediate 

 tergites. Humeral callus with three main setae standing either in a straight 

 line or in a triangle. 9 without outwardly directed prevertical setae and with 

 the usual pair of outer vertical setae. Pre-alar seta very small, shorter than 

 first post ia seta ............ 2 



2 Humeral callus with the three main setae standing in a straight line. Lower calypter 

 with the outer edge distinctly bent downwards. Arista with second segment 

 elongate, about 4-5 times as long as broad. Head with two pairs of strong 

 reclinate orbital setae. Mid tibia with one ad seta. Vibrissae level with 

 epistomal margin ...... ETHILLA Robineau-Desvoidy 



- Humeral callus with the three main setae standing in a distinct triangle. Lower 



calypter not noticeably bent downwards. Arista with second segment not 

 elongate, only about as long as broad. Head without definite reclinate orbital 

 setae (at least in g, probably one pair in $). Mid tibia with two ad setae. 

 Vibrissae inserted well above level of epistomal margin 



MYCTEROMYIELLA Mesnil 



Tribe WINTHEMIINI 



This group is very closely allied to the Carceliini and the Sturmiini and its 

 recognition as a tribe is only doubtfully justified. Townsend (1941) placed Winthemia 

 and its allies in the Sturmiini, but recent workers universally treat the winthemiines 

 as a valid tribe and this course is followed for the present work. The arrangement 

 of the humeral setae in Winthemia is rather distinctive, there being (in males at 

 least) five setae in which three are strong and are spaced in a subtriangular arrange- 

 ment; in this respect Winthemia differs from the Sturmiini in the strict sense in 

 which the humerals are four arranged as three in a straight line and one set forwards. 

 Mesnil (1944) used the presence of five humeral setae as a key character for recognition 

 of the winthemiines, but in reality the character is poor - often in other winthemiine 

 genera than Winthemia itself (e.g. in Nemorilla) five setae may not be definitely 

 developed in males and are typically not developed in females (which usually have 

 four humerals) . 



The winthemiines are best recognized among the Australian Goniinae by the following 

 features in combination (though some Sturmiini and some Carceliini may show some of the 

 features together): eyes densely haired; males without definite reclinate orbital setae; hind 

 tibia (especially in males) with a well developed close-set ad fringe; barette completely haired 

 (or nearly so) ; scutellum with strong crossed horizontal apical setae and with subapical setae 

 inserted widely apart (their bases as distant from each other as base of either from the basal 

 seta of the same side, or even more distantly separated than this) (as in Carcelia, Text-fig. 72). 

 A noteworthy feature of several forms is the presence of pale yellow hair on the presternum 

 instead of the usual black hairs or setulae. 



There is close superficial resemblance between Winthemiini and Ethillini, and 

 as the genus Mycteromyiella (Ethillini) could easily be confused with the winthemiines 

 it is included in the following key. 



Mesnil (1949a : 80) treated Crypsina as a subgenus of Winthemia but it is here 

 preferred to revert to its original generic status. This course maintains Winthemia 

 as a more homogeneous genus, though - certainly - Crypsina is very closely allied. 



