go R. W. CROSSKEY 



2 Eyes bare. Apical scutellar setae very small, directed upwards and either not 



meeting or crossing at tips. Subapical scutellar setae not conspicuously distant 

 from each other, their bases separated by a distance not greater than that 

 between a subapical seta and its corresponding basal seta. Upper occiput 

 with some fine black setulae behind postocular row 



ARGYROPHYLAX Brauer & Bergenstamm 



— Eyes haired. Apical scutellar setae strong, crossed and horizontal. Subapical 



scutellar setae very widely separated, distance between bases much greater 

 than that between base of a subapical seta and its corresponding basal seta 

 (Text-fig. 72). Upper occiput without black setulae behind postocular row . 3 



3 Dorsocentral setae 3+4. Scutellum with extremely strong apical setae that are 



at least as strong as, and usually larger than, the lateral setae. Hairing of 

 eyes long and dense, very conspicuous. Barette normally with hairing, at 

 least at anterior end. Wings hyaline. Not conspicuously black forms . . 4 



— Dorsocentral setae 3 + 3 (four post dc setae in occasional specimens). Scutellum 



with apical setae normally shorter and weaker than lateral setae. Hairing of 

 eyes rather short and usually rather sparse, not so immediately conspicuous. 

 Barette totally bare. Wings suffused with brown anterobasally. Conspicuously 

 black forms with shining abdomen which shows bright silver pollinose areas on 

 each side of T3-T5 (these especially noticeable on T4, at least in §) 



ARGYROTHELAIRA Townsend 



4 Facial ridges bare. Barette when haired with hairing confined to anterior half 



(except sometimes in specimens of Carcelia s. str.) . CARCELIA Robineau-Desvoidy 



- Facial ridges closely setulose up most of their height (Text-fig. 48). Barette 



completely haired CARCEL1MYIA Mesnil 



Key to Australian Subgenera of CARCELIA 



1 Mid tibia with a submedian v seta. Ocellar setae absent or very small, fine and 

 wiry. g with one pair of reclinate orbital setae, $ usually with two pairs (specimens 

 with two such setae on one side and one on the other occur). Hind coxa bare 

 or setulose posterodorsally . . subgenus CARCELIA Robineau-Desvoidy 



- Mid tibia without a v seta. Ocellar setae very strong (except in murina where 



absent or hair-like). Both sexes with two pairs of reclinate orbital setae. Hind 



coxa entirely bare posterodorsally. . . subgenus SENOMETOPIA Macquart 



Tribe ANACAMPTOMYIINI 



Townsend (1940) recognized a tribe for Anacamptomyia and its allies and 

 placed it near Compsilurini (i.e. Blondeliini) , but Mesnil (1944) included the 

 anacamptomyiines among the carceliines. On adult morphology Mesnil's placement 

 seems to be more appropriate than Townsend's, and in fact it is difficult to find 

 characters that satisfactorily differentiate the Anacamptomyiini from the Carceliini. 

 Nevertheless the Anacamptomyiini is here treated as a valid tribe because of its 

 biology: the members of the tribe are unique amongst the Goniinae in being 

 parasites of paper-making wasps (such as Belonogaster and Polistes), and this 

 host-relationship is so exceptional that it appears justified to give it considerable 

 weight in assessing whether or not to treat the flies involved as carceliines or not. 

 It appears possible that the resemblances between Carceliini and Anacamptomyiini 

 are mainly convergent. 



