96 P. E. S. WHALLEY 



(PL 56, figs 335-338), and one or two specimens had a very reduced signum. How- 

 ever no significant trend in the reduction of the size of the signum could be found 

 and no other feature could be found to correlate with this. 



The amount of variability in this species and in the closely allied Madagascan 

 species make it difficult to assess the whole question of speciation in this complex. 

 At present I feel it is not possible to separate significant subspecies in this complex 

 on the morphological evidence available. I regard the whole African-Madagascan 

 complex as a superspecies in which I separate the Madagascan one on certain constant 

 differences from the African species. This superspecies is in the process of different- 

 iating in Africa, while the more isolated Madagascan population has established 

 some phenotypic differences which are constant and presumably now established in 

 the genotype. 



This whole species complex would probably be suitable for a detailed statistical 

 examination on a numerical taxonomic basis. 



Distribution. Map 7 (Groups A and B). 



Group A (strongly bipectinate antennae in male). Senegal; Gambia; Guinea; 

 Ivory Coast ; Ghana ; Togo ; Nigeria ; Central African Republic ; Democratic Republic 

 of the Congo; Uganda; Kenya; Tanzania; Ethiopia; Sudan; Malawi; Zambia; 

 Rhodesia; Angola. 



Group B (very short antennal pectinations in male). Guinea; Nigeria; Democratic 

 Republic of the Congo; Uganda; Kenya. 



Material examined. 



Group A. Holotype <$ (serraticomis) , Uganda: Katagrukwa R., Unyoro, 

 2i.v.[i8]97 (Ansorge), BM slide no. 9520, in BMNH. Holotype £ (cymoeasticha) , 

 Nigeria: 100 ml N. of Lokoja (Cator), BM slide no. 9717, in BMNH. Holotype $ 

 (phricosticha) , Nigeria: 100 ml N. of Lokoja (Cator), BM slide no. 9522, in BMNH. 

 Holotype <$ (nudicornis) , Togo: Bismarckburg, I5.x.[i8]92 (Conradt), in ZMB. 



Senegal: 5 $, Sedhiou, vi-viii.1917 (Castell); Gambia: i $, Bathurst (Carter); 1 <$ 

 (Carter); 1 $ (Moloney); Guinea: 7 <J, 4 $, Beyla (Mrdzek), six <J, four 9 in MMB; 

 Ivory Coast: 6 $ (ex Joicey coll.); Ghana: i $, Gambaga (Bury); 5 <$, 9 <j>, N. 

 Territories, Kete-Krachi (Cardinall); 1 $, Bolgatange, v. 1962 (Lewis); Nigeria: 

 2 % Abinsi, River Benue, vi.1912; 1 $, Ogruga; 1 $, Zungeru, 26.iv.19n (Scott- 

 Macfie); 1 $, Zungeru, 23.X.1910 (Scott-Macfie); 2 $, Zungeru, 28.iii.1911 (Simpson); 

 Central African Republic : 4 <$, 2 $, Fort Crampel (Le Moult) ; Democratic 

 Republic of the Congo : 1 $, Kafakumba, 4.L1925 ; 1 $, Kafakumba, ii.1923 (GFO) ; 

 1 <$, Luvua River (East bank), 85 miles N. of L. Mweru, 3000 ft, end of wet season, 

 iv.1922 (Barns); 2 $, Dungu, Upper Uelle Distr., iv; 4 <$, 1 $, Elisabeth ville, 

 vii.1951-xii.1956 (Seydel), in MNHN; 2 $, Kolwezi, Katanga, x.1960 (Allard); 1 <J, 

 Lulua, Sandoa, iv.1931 (Overlaet), in MRAC; 1 $, Haut-Uele, Motoalrima, xii.1926 

 (Burgeon), in MRAC; Kenya: 2 $, Kibwezi, v. i960 (Carcasson), in NMK; 1 £, 

 Kitale, xii.1961 (Dougall), in NMK; 1 <J, 1 $, Yatta Kitui, iv.1960 (Carcasson); 4 $, 

 Kitale, v.i925-iv.i93i (Jeffery); 1 $, Kitale, vi-viii.1951 (Howard), in NMK; 5 <$, 

 Suna, S. Kavirondo, xi.1931-v.1932 (Feather); Uganda: i $, Madi Opei, Acholi, 

 iii.1952 (Jackson); 1 $, Unyoro, Wakibara, xi.1907 (Ansorge); 1 $, Dokolo, Lango, 



