ioo P. E. S. WHALLEY 



Symphleps seta (Viette) 



Proterozeuxis seta Viette, 1958 : 208. 



Symphleps seta (Viette) Whalley, 1967 : 18, figs 18, 42. 



Distribution. Madagascar. Map 6. 



HAP AN A Whalley 



Hapana Whalley, 1967 : 37. Type-species: Hypolamprus verticalis Warren, by original 

 designation. 



Two of the species [verticalis and carcealis) are very similar in pattern but the 

 third African species (minima) differs in pattern and some other characters and is 

 only tentatively placed in this genus. 



The Madagascan species, H. milloti Viette, although very similar to H. verticalis 

 externally, differs in lacking the tarsal spines. It is interesting that two apparently 

 very closely related species should show this particular difference, in other species 

 in the family differences in the presence or absence of spines on the tarsi are often 

 linked with other fairly major differences. 



This genus can be separated from Hypolamprus by the fusion of R z + R A in the fore 

 wing (these arise from the cell in Hypolamprus). From Rhodoneura it is less clearly 

 separated, the main difference being in the type of signum in the female. Possibly 

 when more species of the genus Rhodoneura are examined the relationship of these 

 two genera may be clearer. 



The genus Hapana is at present known from Madagascar, Aldabra, Mauritius, 

 Africa and North India but possibly other species from the Indo- Australian region 

 should be placed in this genus. The Indian species, Hapana obscuralis Hampson 

 (comb . n .) is very similar to H . verticalis. The males are indistinguishable externally 

 and the genitalia are very similar. There are differences in the female in the shape 

 of the ostium and in the bursa. These two species are closely allied and probably 

 relatively recently separated from a common ancestor. The Aldabran specimens 

 of H. carcealis are more distinct from the mainland specimens of this species than the 

 mainland ones are from specimens from Mauritius. It is possible that the Aldabran 

 specimens should be separated as a distinct subspecies but more information on this 

 species and the allied Madagascan species, H. millotti Viette, is needed. 



Generic description. Eyes without interfacetal hairs. Labial palps 3-segmented. Pro- 

 boscis present. Antennae minutely ciliate. Fore tibia with epiphysis. Hind tibia with two 

 pairs of spurs. Usually one pair of apical tarsal spines. Fore wing with R 3 -\-R t (not R 2 +R t 

 as given in error in the original description, Whalley, 1967 : 37). Uncus simple. Female with 

 signum in bursa. 



Biology. No information. 



Key to the African Species of HAPANA 



1 Very small (6-7 mm wing), reddish brown, small white mark subapically, more 



clearly visible on underside. Male genitalia with long median process on 

 gnathus. Aedeagus with minute spines. Female with twin signa in bursa 



minima (p. 103) 



