106 P. E. S. WHALLEY 



than the other specimens and have the lateral juxta arms more sclerotized. They 

 may represent a distinct subspecies. 



Distribution. Map 41. Guinea; Sierra Leone; Liberia; Ivory Coast; Ghana; 

 Nigeria; Central African Republic; Democratic Republic of the Congo; Uganda; 

 Malawi. 



Material examined. 



Holotype,^ (ramiculata) , Uganda: Masindi, i.i.1898 (Ansorge), BM slide no. 8511, 

 in BMNH. Holotype^ (ansorgei), Uganda: Mondo, 11.iii.18g9 {Ansorge), BM slide 

 no. 9519, in BMNH. 



Guinea: 3 <$, Zoubouroumai, 15 ml. SE. Macenta, 2000 ft, 23. v. 1926 (Collenette); 

 Sierra Leone: 1^,1$, Bo, iii.1967 (Revell); 1^,1$, Bo, iv.1967 (Revell); 1 £, Bo, 

 i-iii.1969 (Revell); Liberia: 2 $, Nimba, Grassfield, vi-vii.1967 (Forbes-Watson); 

 Ivory Coast: i $, Aedguie, ix.1964 (Griveaud), in MNHN; Ghana: i <$, Winnebah, 

 30.1.1940, in CMP; 1^,1$, N. Territories, Kete-Krachi (Cardinall); Nigeria 

 2 (J, 1 $, Sapoba, Benin, ex Terminalia ivoriensis, ix.1962; 1 $, Sapoba, ex Termi- 

 nalia ivoriensis, vii.1968; 1 $, Ife-Ikeji, ex Terminalia ivoriensis vi.1968; Central 

 African Republic; i (J, Fort Crampel (Le Moult); Democratic Republic of 

 the Congo: i $, Dungu, Upper Uelle Distr., vii. ; 1 <$, Lulua, Kapanga, 1933 

 (Overlaet), in MRAC; Uganda: 3^, Mabira Forest, Jinja, x.1962 (Carcasson), two 

 cj in NMK; Malawi: i $, Zomba Plateau, 1920 (Barlow). 



OPULINI 



This tribe is separated from the Rhodoneurini by the absence of spines on the 

 tarsal segments (see page 84). In a few species, spines may be present on the last 

 tarsal segment of the hind leg. 



EPAENA Karsch 



Epaena Karsch, 1900 : 245. Type-species, Epaena elephantinalis Karsch, by monotypy. 

 [Tridesmodes auct., nee Warren.] 

 Epaena Karsch, Whalley, 1964a : 120. 



This genus, which contains eight species, is divided into three species-groups. 



This whole genus is rather heterogeneous and may well be further subdivided 

 when more specimens are studied and more information about species in other 

 faunae is available. The genitalia of E. trijuncta are slightly different from the rest 

 of the species in the genus. At present the species of the genus are only known 

 from the mainland of Africa. Epaena possesses certain characters which make it a 

 possible link in the Rhodoneura-Tridesmodes group of genera. Epaena differs from 

 both these genera in lacking tarsal spines but it does have the same veins fused in the 

 fore wing as Tridesmodes and a similar shaped gnathus and general wing colour and 

 pattern to some species of Rhodoneura. 



Generic description. Labial palps 3-segmented. Eyes without interfacetal hairs. Pro- 

 boscis present. Antennae, ciliate, minutely ciliate or bipectinate. Fore tibia with epiphysis. 

 Hind tibia with two pairs of spurs. Tarsi without spines. Fore wing usually with some fusion 



