THYRIDIDAE OF AFRICA AND ITS ISLANDS 109 



- Fore wing without obvious black marks on hind margin. Genitalia not as above 2 

 2 Valve of male broad, gnathus with minute spines .... inops (p. 109) 



- Valve of male narrowing near apex. Gnathus without spines . Candida (p. no) 



Epaena inops (Gaede) comb. n. 



(PI. ii, fig. 51; PI. 35, fig. 194; PI. 58, figs 352, 353) 



Symphleps inops Gaede, 1917 : 382. 

 Symphleps inops Gaede; Gaede, 1929 : 498. 



(J. Wing, 12-5-21 mm. Vertex brown, frons bulbous with tuft of scales protruding slightly 

 between eyes. Labial palps with third segment more than 1/2 length of second, upturned, 

 just reaching vertex. Patagia brown. Thorax white, irrorate with brown. Hind tibia with 

 inner spur of distal pair long, more than 2 X length of outer spur. Inner spur slightly more than 

 1/2 length of 1st hind tarsal segment. Fore wing, pattern as in PI. n, fig. 51, white with brown 

 reticulations (white phase). Underside reticulations darker with brown costal, terminal and 

 apical margins. Veins R 3 and R t anastomosing, arising from common stem of R 2 +R 3 +R t . 

 Veins lA and iA join near base to form single vein to wing margin. Hind wing, colour and 

 pattern as fore wing. Sc+R t and Rs approach but do not join. 



Genitalia 3 (PI. 35, fig. 194). Uncus simple. Gnathus with median process covered with 

 small spines. Valve simple, basal process sclerotized, slightly hooked. Juxta two rounded 

 lobes with broader plate at 90 to them. Aedeagus with spiny vesica and small spines on 

 manica. 



$. Wing, 20-24 mm. Colour and pattern as in male. Labial palps with third segment 1/3 

 length of second. Venation as male. 



Genitalia $ (PI. 58, figs 352, 353). Anal papillae short. Ostium and duct slightly enlarged 

 and sclerotized. Duct spiny. 



Discussion. This species occurs in two colour phases. One phase is completely 

 white with brown pattern, the other is brown with a darker brown pattern. The 

 genitalia of these two phases are similar and intermediates between the two 

 extremes occur. The pattern is variable but the reticulations are generally paler 

 than in the other white African Thyridids. This species also varies in size and in the 

 intensity of pattern and will probably be separated into subspecies when more 

 material is available. The Angolan and Cameroon specimens are smaller and with 

 more intense pattern than the Rhodesian or Congo ones but the genitalia are similar. 

 Unlike some of the other species in the genus, no variation in the origin of the radial 

 veins was found. 



Distribution. Map 38. Cameroon; Democratic Republic of the Congo; 

 Tanzania; Zambia; Angola. 



Material examined. 



Holotype <$, Tanzania: Udjidji, BM slide no. 9676, in ZMB. Cameroon: i $, 

 Efulen (Weber), v.1917, in CMP; 1 <$, Bitje, Ja River, 200 ft, x-xi.1912; Democratic 

 Republic of the Congo: i<£, i $, Katanga, Kolwezi, x.1954 (Allard) ; 1 <J, Katanga, 

 Tenke, vii-viii.1931 (Cockerell); 1 <$, Katanga, Jadotville, viii.1965 (Coussement); 

 Zambia: i <$, Abercorn, xi.1963 (Vesey-Fitzgerald) , in NMK; 1 $, Chingola, ii.1960, 

 in NMR; 1 <$, Mpiko, iii-vii.1921 ; 2 $, 1 ?, Kitwe, 2-27. xii. 1954 (Kruger), one £, 

 one $, in TMP; 1 <J, 1 $, Ndola, xii. i960, in NMR; Angola: i £, Gamba, Bihe, 

 xii. 1934 (Braun); Andulo, Bihe, xii. 1934 (Braun). 



