n 4 P- E. S. WHALLEY 



ville and Coquilhatville along the River Congo], xi.1921 (Verlaine), BM slide no. 

 9939, in MRAC. 



Paratype. Democratic Republic of the Congo: i $, Opala, Lomami R., 

 Prov. Orientale, iii.1959 (Carcasson). 



Epaena radiata (Warren) comb. n. 



(PI. 2, T; PL 36, fig. 200; PI. 59, figs 358, 359) 



Beguma radiata Warren, 1908 : 328. 

 Beguma radiata Warren; Dalle Torre, 1914 : 39. 

 Beguma radiata Warren; Gaede, 1917 : 358. 

 Beguma radiata Warren; Gaede, 1929 : 498. 



o*. Wing, I2-5-I5-5 mm. Vertex brown, frons rounded, produced between eyes. Antennae 

 bipectinate. Labial palps short, slightly less than diameter of eye, third segment 1/3 length 

 of second. Thorax dark brown. Hind tibia with proximal pair of spurs short, distal pair with 

 outer spur slightly shorter than inner spur, longest spur less than 1/2 length of 1st hind tarsal 

 segment. Tarsi mostly without spines, some spines on last tarsal segment on hind leg. Fore 

 wing, pattern as in PI. 2, T, dark chocolate-brown with lighter and darker transverse fasciae. 

 Frenulum single. Underside paler. Vein R 3 and i? 4 shortly stalked. 2A reduced to short 

 stalk off \A . Hind wing, pattern as fore wing, Sc +Rx and Rs free. 



Genitalia <J (PI. 36, fig. 200). Uncus broad. Gnathus a sclerotized loop without median 

 process. Valve small, narrowing in apical third. Juxta lightly sclerotized. Basal process 

 of valve a broad plate, lightly sclerotized. Aedeagus with few minute spines on vesica. 



$. Wing, 16 mm. Paler coloured than male, pattern similar. Labial palps i-i£ X 

 diameter of eye. Frenulum with 4-5 bristles. 



Genitalia $ (PI. 59, figs 358, 359). Anal papillae short. Ostium broad, first part of duct 

 looped, remainder broad, no signum. 



Discussion. The pattern of this species immediately separates it from all 

 other African Thyrididae. It varies in intensity of colour but there seems to be little 

 variation in pattern. This species also has a number of morphological peculiarities. 

 The position of this species within the genus Epaena is uncertain and it may need a 

 separate genus. There may be some related Indo- Australian species but the wing 

 pattern of radiata is unique in the family. With the morphological differences, it is 

 not possible to suggest which species are most closely allied to radiata but it fits 

 broadly into the generic concept of Epaena given on page 106. 



Distribution. Map 40. Guinea; Liberia; Ivory Coast; Ghana; Nigeria; 

 Gabon; Democratic Republic of the Congo; Uganda; Sudan. 



Material examined. 



Holotype^, Democratic Republic of the Congo; Luebo, Kassai R. (Landbeck), 

 BM slide no. 9612, in BMNH. 



Guinea: i $, Soundedou, nr Macenta, 1600 ft (Collenette), v. 1926, at light; 

 Liberia: i <$, Harbel, Marshall Terr., 14. v. 1956 {Fox), in CMP; Ghana: i <$, 

 Bibianaha, 70 miles NW. of Dimkwa, 700 ft (Spurrell); Ivory Coast: 1^,1$, 

 Bingerville [Melon), vi.1915; 1 $, 1 $, Foret du Banco, ix.1963 (Piart & Griveaud), in 

 MNHN; 1 $ (Oberthur coll.) ; Nigeria: i (J, Ibadan, Jericho, 3.1V. 1959, at light; 2 <$, 

 Ibadan (Bowden); 1 $, Lagos (Strachan); Gabon: i <$, Belinga (Grasse), Miss. Biol. 

 Gabon, in MNHN; Democratic Republic of the Congo: i <$, Ifuta, x.1921; 1 <J, 



