12 W. A. SANDS 



(8) Pilosity of imago head capsule sparse, fine or inconspicuous, or more 



numerous and coarser 



(9) 'Abdominal dehiscence' in worker caste present or absent 



(10) Mixed segment of worker gut with or without inflated section at distal end 

 (11-14) Enteric valve of worker gut: sclerotization of cushions present or absent 



(each of four independent positions) 

 (15-18) Enteric valve : position of spines of main armature on cushions, proximal 



third, present or absent. ('Main armature' refers to scales or spines at 



least one place higher in rank under characters 88-91 than remainder. 



If spines uniform then all scored zero here) 

 (19-22) Enteric valve: spines of main armature present or absent in mesal third 

 (23-26) Enteric valve : spines of main armature present or absent in distal third 



(27) Enteric valve: membranous wall between and beyond cushions smooth, 



or carunculate or scaly 



(ii) Multi-state qualitative 



(28) Fontanelle of imago: shape; circular, short oval or elongated 



(29) Fontanelle of imago: level; depressed, flat, or slightly raised 



(30) Fontanelle of imago : size ; vestigial, less than ocellus or more than ocellus 



(31) Medial spot of imago (i.e., small spot just anterior to fontanelle) ; shape (as 



above) 



(32) Medial spot of imago : level as above 



(33) Medial spot of imago: size; less than, equal to, or more than fontanelle 



(34) Frontal marks of imago (between ocelli and postclypeus) ; shape, indistinct 



or vestigial, crescent-shaped, or semicircular 



Quantitative : 



(i) Continuous variables (measurements) 



(35) Imago : width of head capsule across compound eyes (W) 



(36) Imago: least diameter of ocellus (O w ) 



(37) Imago : greatest diameter of ocellus (Oi) 



(38) Imago : distance ocellus to compound eye (O-E) 



(39) Imago : length of postclypeus (maximum) (Pel) 



(40) Imago: length of antennal article 3 (III) (Text-fig. 1) 



(41) Imago: length of antennal article 4 (IV) 



(42) Imago: left mandible, distance from apical to first marginal tooth (La) 



(Text-fig. 2) 



(43) Imago: left mandible, distance from first to third marginal (Li) 



(44) Imago: left mandible, distance from third marginal to end of subsidiary 



tooth (L m ) 



(45) Imago: right mandible, distance from apical to first marginal tooth (Ra) 



(Text-fig. 3) 



(46) Imago: right mandible, distance from first to second marginal (Ri) 



(47) Imago: right mandible, distance from second marginal to notch between 



it and the molar plate (R m ) 



