128 W. A. SANDS 



setae. In A. brevior and A. quietus, the right first marginal is reduced, and shorter 

 than the second. In A. empodius it is the subsidiary marginal tooth of the left 

 mandible which is scarcely clear of the molar prominence; the fontanelle is pale and 

 depressed. A. irrixosus is again very similar to Alyscotermes and was at first placed 

 in that genus. The absence of enteric valve armature is in this case the only 

 distinguishing feature. 



Clearly Astalotermes and Alyscotermes are closely related, and, as indicated in the 

 discussion on Astalotermes, other ways of delimiting genera to take account of this 

 have been considered. The results of the principal co-ordinate analysis separate 

 them, as do the cluster analyses. In order to be consistent with the results for 

 some better defined genera, they are named as separate taxa. In the numerical 

 analyses, Alyscotermes is placed nearest to Ateuchotermes. (Text-figs 304, 305 & 

 306 to be compared with Text-figs 400, 401 & 402.) The latter genus has a dis- 

 tinctive enteric valve and no confusion can arise between them. 



The name of this genus refers to the frequency with which it has displayed the 

 behaviour described by Grasse" & Noirot (1951) as 'La sociotomie'. Entire colonies 

 have been found migrating complete with physogastric queen over the soil-surface. 

 This usually results from an attack by burrowing Doryline ants. The termites flee 

 from their beleaguered nest tunnels and wander about looking for a new home. 



No key is provided since the genus contains only two species, and presents no 

 problem of identification once the generic identity of a specimen is recognized. 

 A. kilimandjaricus (Sjostedt) is widespread and common, A. trestus sp. n. is only 

 known from one nest-series. It is distinguished from its commoner congener by 

 the much more spiny enteric valve armature and slightly differently proportioned 

 mandibles of the worker caste. The only difference in the imago of A. trestus is 

 the more evenly rounded postclypeus. 



Alyscotermes kilimandjaricus (Sjostedt) comb. n. 

 (Text-figs 307-310 & 315-320; PI. 2, figs 8-1 1) 



Entermes kilimandjaricus Sjostedt, 1907 : 9. LECTOTYPE $, Tanzania: Kilimandjaro, 



Kibonoto (Naturhistoriska Riksmuseum, Stockholm), here designated [examined]. 

 Mirotermes {Cubitermes) natalensis Holmgren, 1913 : 355. Type-series, Republic of South 



Africa: Natal, Amanzimtoti (1 $ syntype [examined] in American Museum of Natural 



History; rest of type-series in Mus. Gotenborg). Syn. n. 

 Mirotermes (? Procubitermes) mfolozii Fuller, 1925 : 190. LECTOTYPE q\ Republic of 



South Africa: Zululand, White Mfolosi River, Conjeni (National Collection of Isoptera, 



Pretoria), here designated [examined]. Syn. n. 



Imago. Head capsule sepia to very dark sepia or chestnut-brown; sometimes darker above 

 ocelli; fontanelle broad to somewhat elongate oval, flat, ridged in middle or slightly raised 

 smaller to distinctly larger than ocellus, orange-yellow to dark sepia-brown; medial spot flat 

 or slightly raised, oval, smaller than fontanelle, coloured as head or a little paler; postclypeus 

 yellow-brown to sepia-brown, labrum yellow to brown ; frontal marks flat or slightly depressed 

 crescents, distinctly paler than head; antennae yellow-brown to sepia-brown. Pronotum 

 brown to dark sepia, meso- and metanota brown, transverse dark sutures variable, weak to 

 distinct; femora pale yellow-brown to brown, tibiae paler, tarsi yellow. Abdominal tergites 



