i3o W. A. SANDS 



Right mandible, first to second 



marginal (Ri) . . . 0-11-0-15 0129 ± 0009 



Right mandible, second marginal 



to molar (R m ) ■ • • 0-08-0-10 0082 i 0-006 



Mesonotum width (M) . . 0-30-0-44 0365 ± 0-036 



Metanotum width (N) . . 0-30-0-43 0369 ± 0-037 



Worker. Head capsule yellow-white to yellow, pilosity yellow, sparse. Postclypeus weakly 

 to moderately inflated, Pcl/W, 0-21-0-29, Pcl/Ri, 1-89-2-71. Left mandible with subsidiary 

 marginal tooth about same size as third marginal, separated from molar prominence by wide 

 deep notch in surface view, complex ratio, LA/Li-L m , 7-90-12-75, anterior edges of first and 

 second marginal teeth of right mandible equal, Ra/Ri-Rmi. 10-49-15-40. Enteric valve armature 

 in cushion position 3 with 1-9 (only two recorded with more than five) pronounced spines, 

 position 4 sometimes not produced through valve opening, rarely with more than one pronounced 

 spine of ten only small spines, both positions with further small or minute spines; positions 1 

 and 2 with a few small spines, or all spines minute. Membranous wall of valve beyond cushions 

 with minute spicules. Other characters as in generic diagnosis. 



Measurements (in millimetres). 



' A. kilimandjaricus' 'A. iiatalensis' 



Range Mean ± S.D. Range Mean ± S.D. 



Head width (W) .... 0-81-1-05 °'934 ± 0069 0-84-1-00 0-925 ±0-051 



Fore tibia width (T w ) . . . 0-09-0-14 0-114^0-010 0-11-0-13 0-115 ±0007 



Fore tibia length (Tj) . . . 0-56-0-84 0-689 i 0-090 0-63-0-71 0-678 ± 0-030 



Postclypeus length (Pel) . . 0-19-0-33 0-268^0-032 0-21-0-26 0-235^0-017 



Left mandible, apical to first marginal 



(La) ...... 0-08-0-10 0088 Jz 0-005 0-08-0-09 0087 i 0-006 



Left mandible, first to third marginal 



(Li) ...... 0-13-0-16 0148 4^ 0010 013-015 0143 Az 0007 



Left mandible, third marginal to 



molar (L m ) . .... 0-05-0-07 0-060^0-007 005-006 0058^0-005 



Right mandible, apical to first mar- 

 ginal (Ra) ..... 0-08-0-09 0-087 ± 0-005 0-07-0-09 0-083 ± 0-006 



Right mandible, first to second mar- 

 ginal (Ri) ..... 0-09-0-12 0109 ± 0008 009-011 0-103^0-006 



Right mandible, second marginal to 



molar (R m ) ..... 0-05-0-07 0-060 i 0-005 0-05-0-06 0060 x 0-004 



The essential differences between A. kilimandjaricus and A. trestus have been 

 mentioned under the generic heading in lieu of a key. The two are clearly very 

 closely related, and A. trestus appears to be a local species that has arisen in the 

 anomalous and ecologically isolated area of the Mara forest. A. kilimandjaricus 

 has the widest distribution of any of the species in this monograph. It is found 

 from the Gambia in West Africa to Cape Province in South Africa, and in conse- 

 quence has one of the longest ranges of any African termite. Until a late stage 

 in the present work it was regarded as two species, A . kilimandjaricus for the northern 

 specimens and A. natalensis for the southern. Slight differences in shape, pilosity 

 and in the gut-characters were thought to occur between them, and were coded 

 accordingly for their separate inclusion in the similarity analysis. However, 

 re-assessment of these features took place when it became necessary to key out the 

 two species. No clear distinction could be made between them on any character 

 or combination of characters and there are not sufficient differences even to warrant 



