PTEROMALIDAE OF N.W. EUROPE 633 



Type material. Holotype $ and allotype <$ in Narodni Museum, Prague (Cat. 

 nos. 3053, 3054) ; paratypes in Forest Research Institute, Warsaw ; Poland, 

 Pieniny, Polish Carpathians, reared in 1948 from Pityophthorus polonicus Karp., 

 (/. /. Karpinski). 



Norway, Sweden, Finland, Poland. 



Biology. Parasite of Coleoptera Scolytidae, e.g., Pityophthorus polonicus Karp. 

 (Boucek, 1954a), Pityogenes quadridens (Htd.) Bakke, 1956 ; Hedqvist, 1963), 

 Pityogenes chalcographus (L.) (Nuorteva, 1957 ; Hedqvist, 1963). 



EUMACEPOLUS Graham 



Eumacepolus Graham, 1957c : 137-138. Type-species : Pteromalus saxeseni (Ratzeburg) sensu 

 Thomson, 1878, by monotypy and original designation [= Eumacepolus grahami v. Rosen, 

 i960]. 



Eumacepolus Graham ; v. Rosen, 1960a : 48. 



Eumacepolus Graham, 1961 : 173-176. 



When the genus was described, the type-species was designated as " Pteromalus 

 saxeseni (Ratzeburg) Thomson (=Habrocytus saxeseni (Ratz.) Thomson, 1878) ". 

 My reason for specifying that the type-species was saxeseni Ratzeburg in the sense 

 of Thomson was that I felt some doubts as to whether Thomson had correctly 

 interpreted Ratzeburg's species. My action proved to be well-founded because 

 von Rosen (1959 : 134-6) showed that Thomson's interpretation of Pteromalus 

 saxesenii Ratzeburg was in fact incorrect ; consequently (1960a : 48) he renamed the 

 type-species of Eumacepolus as E. grahami v. Rosen. Because of the way in which 



1 originally formulated my designation of the type-species, the genus name remains 

 valid. 



The mandibles are exceptionally variable in this genus. In pulcher, both 

 mandibles may have 4 teeth ; or the left mandible may have 3 and the right one 4. 

 In the specimens of obscurior so far captured, both mandibles have 3 teeth (the 

 inner tooth being broadly truncate in both mandibles). In grahami the mandibular 

 formula is 3.4. 



The female recorded by me (1961 : 175) as being probably that of obscurior, does 

 not belong to it but evidently represents an undescribed species. The characters of 

 what is almost certainly the true female of obscurior are given in the accompanying 

 key. 



Eumacepolus presents some taxonomic problems. The species grahami differs in 

 several respects from the others (see key) and appears to be very close to 

 Mesopolobus ; it differs from all our species of Mesopolobus, however, in having only 



2 anelli followed by a relatively elongate first funicular segment. In Mesopolobus, 

 when only two anelli are present, the first funicular segment is relatively short, 

 subquadrate to transverse, and often shorter than the second funicular segment. 



On the other hand, Eumacepolus pulcher and E. obscurior appear to be less close 

 to Mesopolobus and to resemble in some ways certain species of Ablaxia ; they 

 seem more appropriately placed in a distinct subgenus of Eumacepolus. 



